Bulletin of the

British Museum (Natural History)

(NAMl HISTGSY)

- 1AUG'i988

PRESENTED
GENER/ 1 ^RY

An illustrated catalogue of the type
specimens in the Greville diatom
herbarium

David M. Williams

Botany series Vol 18 No 1

28 July 1988

The Bulletin of the British Museum (Natural History], instituted in 1949, is issued in four
scientific series, Botany, Entomology, Geology (incorporating Mineralogy) and Zoology,
and an Historical series.

Papers in the Bulletin are primarily the results of research carried out on the unique and
ever-growing collections of the Museum, both by the scientific staff of the Museum and by
specialists from elsewhere who make use of the Museum's resources. Many of the papers are
works of reference that will remain indispensable for years to come.

Parts are published at irregular intervals as they become ready, each is complete in itself,
available separately, and individually priced. Volumes contain about 300 pages and several
volumes may appear within a calendar year. Subscriptions may be placed for one or more of
the series on either an Annual or Per Volume basis. Prices vary according to the contents of
the individual parts. Orders and enquiries should be sent to:

Publications Sales,

British Museum (Natural History),
Cromwell Road,

London SW75BD,
England.

World List abbreviation: Bull. Br. Mus. nat. Hist. (Bot.)

British Museum (Natural History), 1988

The Botany series is edited in the Museum's Department of Botany

Keeper of Botany: Mr J. F. M. Cannon

Editor of Bulletin: Mr J. R. Laundon

Assistant Editors: Dr A. J. Harrington and Miss M. J. Short

ISBN 565 08019 9
ISSN 0068-2292

British Museum (Natural History)
Cromwell Road
London SW7 5BD

Botany series
Voll8Nolppl-148

Issued 28 July 1988

;1AU

An illustrated catalogue of the type specimens in the
Greville diatom herbarium

David M. Williams

Department of Botany, British Museum (Natural History), Cromwell Road, London
SW7 5BD

Contents

Synopsis 1

Introduction 1

Methods 2

Specimen citation 2

Typification 2

Light microscopy 3

Data collection, management, and storage 3

Greville types 3

Appendix 61

Acknowledgements 62

References and bibliography 62

Index 66

List of plates 72

Plates : 75

Synopsis

An illustrated catalogue of the type specimens in Greville's herbarium, which were used to illustrate his
published works, has been prepared. The catalogue has been arranged alphabetically by genus and species
using Greville's original name, regardless of current nomenclatural status, to avoid confusion with current
systems of classification that may differ from each other. 494 taxa are discussed. An index has been
provided that gives a guide to currently accepted names. In cases where the currently accepted names may
be the subject of some debate, a note to that effect is appended to the genus or species under consideration.
It has not been the purpose of this catalogue to rectify any taxonomic inaccuracies, but to typify Greville's
names allowing specialists to make any necessary taxonomic revisions. In some cases it has been necessary
to introduce nomenclatural changes where clear instances of priority have been established.

The following nomenclatural changes have been included in an appendix: Azpeitia obscurum (Grev.) P.
A. Sims comb. nov. (Tricertium obscurum Grev.), Odontella hastata (Grev.) J. Fenner ex D. M. Williams
comb. nov. (Hemiaulus hastatus Grev.), Perissonoe parvula (Grev.) D. M. Williams comb. nov.
(Amphitetras parvula Grev.), and Pseudauliscus grevillei R. Ross nom. nov. (Auliscus nebulosus Grev.).

Introduction

The Greville diatom herbarium consists of 5248 glass microscope slides and a number of
herbarium sheets with packets containing material dried on to mica. The earlier numbers in the
slide collection are made from material Greville received from other phycologists; it contains a
considerable amount of type material published during the early period of diatom taxonomy.
However, the later numbers of Greville's slides are of greater interest as there are very few
duplicates in other herbaria. The herbarium sheet material contains the specimens referred to in
Greville's first diatom publications.

This catalogue has been prepared from the material Greville used to illustrate his own diatom
publications (see bibliography), the most significant of which were twenty papers dealing with
various Recent and fossil localities. Of special importance are the first accounts of diatoms in a
Late Eocene/Oligocene deposit from Barbados, using material received from Christopher
Johnson (Greville, 1861-1866, collected together in Simonsen, 1968). Greville's publications

Bull. Br. Mus. not. Hist. (Bot.) 18 (1): 1-148 Issued 28 July 1988

2 D. M. WILLIAMS

are amongst the earliest for fossil diatom studies, and are of major nomenclatural importance as
they contain descriptions of many species and genera.

The catalogue has been arranged alphabetically by genus and species using Greville's original
name, regardless of current nomenclatural status. This is to avoid confusion with current
systems of classification that may differ from each other. Altogether 494 taxa are dealt with. An
index has been provided that gives a guide to currently accepted names. In cases where these
may be the subject of some debate, a note to that effect is appended to the genus or species under
consideration. It has not been the purpose of this catalogue to rectify any taxonomic inaccur-
acies, but to typify Greville's names, allowing specialists to make the necessary taxonomic
revisions. In some cases it has been necessary to introduce nomenclatural changes where clear
instances of priority have been established. Additionally, some taxa have been revised by
Mr R. Ross but never published.

Most of the specimens Greville described as new have remained in his herbarium. However,
there are exceptions where Greville was either loaned slides or used examples from other
collectors. As a consequence a number of holotype specimens are lodged within other
collections. Fortunately, most of these types have since been donated to the BM.

Methods
Specimen citation

Each taxon has been given its bibliographic citation, indicating the original publication. In a few
cases, when a drawing was published later than the description, both references are given. When
a number of Greville's papers were simultaneously published in two journals, reference has
been given to both. Greville did not publish all the taxa he considered new to science and later
workers used his specimens and names. These taxa appear in Rattray (18880, b, 1890) and Ralfs
(in Pritchard, 1861). As these form part of Greville's collection, they are included in this
catalogue.

The locality data has been transcribed exactly as it appears on the slide label. In cases of
obvious confusion, or where additional information is needed to clarify the meaning, this has
been supplied enclosed in square brackets. In most cases the data on the slide label reflects the
idiosyncrasies of abbreviation or spelling.

Typification

The majority of the glass slides are of strewn material mounted in Canada Balsam. A few
selected mounts Greville received from other collectors and mounters are also present. The first
3000 slides, including those received from other workers, have been fully labelled with the
noteworthy species, whilst the remaining 2000 slides are considered as additional material.
However, in several instances Greville finally chose specimens from these later slides for his
published illustration.

Greville located his specimens either by using a Maltwood Finder (abbreviated as MF in this
work) or by circling the coverglass with a diamond marker around the particular specimen. The
MF is based on a grid mounted on to a glass slide which has two sets of numbers, correlating to
the lateral and vertical movement of the microscope stage. This is not a common method of
specimen location today and few Maltwood Finder's remain in existence.

In addition to the slides, there is a large quantity of manuscript material, including notes,
some copies of submitted manuscripts, and most of the original drawings, preserved in the BM.
The drawings are exceptionally important when deciding on the type status of each specimen.
Greville included on the pencil drawing the slide number and the MF co-ordinates for the
specimen he had under consideration. In most cases, this allows the exact specimen to be
re-located under the microscope. These specimens are referred to as holotypes. In cases where
there is either no drawing or the drawing has either ambiguous or no slide annotations, then all
possible specimens indicated by Greville are referred to as syntypes. Where it has not been
possible to establish the holotype with certainty, a lectotype has been designated.

GREVILLE DIATOM TYPES 3

The terminology of botanical nomenclature was taken from McVaugh, Ross & Stafleu (1968) .
Their definitions of types were adopted. In addition, the term protologue has been used to mean
'everything associated with a name at its first publication' (McVaugh, Ross & Stafleu, 1968: 22).

Light microscopy

A Zeiss microscope, equipped with Leitz objectives and Zeiss eye-pieces, was used throughout
this work. The photographs were taken using an Olympus OM-2 loaded with Ilford FP4, ASA
135 film. All specimens were photographed under oil immersion and phase contrast using the
x 100 objective. If the condition of the slide prevented the use of high magnification, a x 40 dry,
phase contrast objective was used.

Data collection, management, and storage

As information was drawn from several sources, a computer system was used to collate the data.
This resulted in a database containing information from the slide labels, the various manuscript
notes and drawings, and from the published literature. Production of this catalogue has utilized
only a part of the information now stored. The full database is in active use for further projects.
Initially a DMS microcomputer was used but the largest part of the project was completed on
Torchnet equipment. The system uses CP/M operating system with the DMS (Data Manage- <;
ment System) software.

|i
< ?

Greville types
I. ACHNANTHESBory

1. A. angustata Grev. in Q. Jl microsc. Sci. 7: 163, fig. 9 (1859). (PI. 1, Fig. 1; specimen length
130 /am)

Patos guano /. T. Norman 8.1858, BM 1754, holotype.

There is no drawing of this species in Greville's manuscript collection. However, this is the only
slide Greville indicates for this species. The specimen is ringed and shows two frustules that
compare favourably with the published illustration. It is reasonable to assume that this is the
holotype specimen.

2. A. carmichaelii Grev. in Hook., Engl. FL: 404 (1833). (PI. 1, Figs 2-4; specimen length
75 /Jim; specimen width 3 /am in Fig. 3; width 7 /am in Fig. 4)

Parasitic on smaller filiforme algae [Appin, Scotland], Captain Carmichael, BM 81228, BM
81229, BM 81230.

This material forms part of the collection that is on permanent loan to the BM from Kew. Two
slides are mounted in Mikrops 163 and the third is in Naphrax. As the material is from dried
specimens it is difficult to find complete, undamaged valve views. These specimens are syntypes.
The micrographs are taken from BM 81228.

3. A. costata Grev. in Trans, hot. Soc. Edinb. 8: 438, figs 8-10 (1866). (PI. 1, Figs 8-10;
specimen length 37-5 /urn in Fig. 8; 50 /am in Fig. 9; 52 /urn in Fig. 10)

Sandwich Islds, BM 3550, lectotype, BM 3546, BM 3551; Sandwich Islds Prof. H. Smith, BM
3552; Sandwich Isles, BM 3930; Sandwich Islands, BM 3931.

There are no manuscript notes or drawings for this species but the protologue indicates that it is
present in the same gathering as A. pennaeformis Grev. All slides from the type locality were
examined and, as BM 3550 contained three specimens corresponding to the raphe valve, the
raphe-less valve, and the girdle view of the original published figures, it was chosen as lectotype.
The name is a later homonym of A. costata Johnston 1860 and therefore illegitimate.

4. A. gregoriana Grev. in Q. Jl microsc. Sci. 1: 84, figs 13, 14 (1859). (PI. 1, Figs 11-12;
specimen length 175 /am)

Arran 57, BM 2231, lectotype; Arran 1856, BM 3291; Arran 1857, BM 3487; Arran 57, BM
5149.

4 D. M. WILLIAMS

There are four slides in the collection with specimens of A. gregoriana marked by Greville, but
there is no manuscript evidence to indicate which of these sides was used for his type
illustrations. The most representative, BM 2231, specimen no. 2, has been chosen as the
lectotype. There are two specimens on BM 3487, two specimens on BM 3291, and a single
specimen on BM 5149.

5. A. pennaeformis Grev. in Trans, hot. Soc. Edinb. 8: 438, figs 11-13 (1866). (PI. 1, Figs 5-7;

specimen length 140 /xm in Fig. 5; 140 /xm in Fig. 6; 137-5 /am in Fig. 7)

Sandwich Islds, BM 3546, BM 3550, BM 3551; Sandwich Islds Prof. H. Smith, BM 3552;

Sandwich Isles, BM 3930; Sandwich Islands, BM 3931, syntypes.

There is no drawing in the Greville manuscript collection but there are a series of manuscript

notes dealing with this species. On the card cover to these notes, Greville indicates the

specimens that he consulted. These were BM 3546 (MF 26/21; not illustrated), BM 3550 (MF

34/28, 21/27, and 36/26), and he also erroneously refers to BM 3552 (MF 31/32). All these have

been matched to the drawings but it is difficult to establish exact correspondence with certainty.

As Greville included three specimens in his protologue, they are all considered syntypes.

II. ACTINODISCUSGrev.

1. A. barbadensis Grev. in Trans, microsc. Soc. Lond. 11: 69, fig. 11 (1863). (PI. 2, Fig. 1;

specimen diameter 85 /xm)

Barbados* Johnson 1862, BM 2829, holotype.

III. ACTINOPTYCHUS Ehrenb .

1. A. minutus Grev. in Trans, microsc. Soc. Lond. 14: 5, fig. 12 (1866).

'Monterey deposit; cabinet of L. Hardman, Esq. ; very rare'. (Slide not found).

Although there is a drawing of this species (annotated with 'H. 53') in the Greville manuscript

collection, the holotype slide has not been located. It is a Hardman slide, suggested by the 'H'

part of the annotation.

IV. AMPHIPRORA Kiitz., non Ehrenb.

At present there is some controversy concerning the correct nomenclature of this genus
(Paddock & Sims, 1980: 181; see also Ross in Hartley, 1986: 596). Several of Greville's species
have been re-examined and placed in new genera (Paddock & Sims, 1980; Paddock, in prep.). A
revision of the genus, including the Greville species, is in progress (Paddock & Sims, in prep.).

1. A. brebissoniana Grev. in Edinb. Newphil. J. 18: 185, fig. 8 (1863); in Trans, hot. Soc. Edinb.
7: 578, fig. 8 (1863). (PI. 2, Figs 2, 4, 5; specimen length 62-5 /xm)

Curteis Straits Queensland Dr Roberts 1862, BM 2601, holotype.

There is a drawing of this species in the Greville manuscript collection which is annotated with
the numbers BM 2601 (MF 28/16) and BM 2624. On BM 2624 there is a specimen indicated at
MF 31/20, but as this is not specifically referred to on the manuscript drawing, specimen MF
28/16 on BM 2601 must be the holotype.

2. A. chinensis Grev. in Ann. Mag. nat. Hist. HI, 16: 7, fig. 13 (1865). (PI. 2, Fig. 6; specimen
length 113/xm)

Hong Kong 16, BM 4495; Hong Kong, BM 4501, syntype.

The drawing in the Greville manuscript collection, is annotated with 'no. 16'. This is part of a
series of working slides from the Hong Kong material. The only specimens of this taxon to be
discovered in Greville's slide collection are on slide BM 4501. The micrograph is from specimen
MF 25/24.

3. A? clepsydra Grev. in Edinb. Newphil. J. 18: 41, fig. 20 (1863); in Trans, hot. Soc. Edinb. 7:
541, fig. 20 (1863). (PI. 2, Fig. 8; specimen length 70 /am)

Curteis Straits, BM 2612, holotype.

GREVILLE DIATOM TYPES 5

4. A. conspicua Grev. in Trans, microsc. Soc. Lond. 9: 86, fig. 16 (1861). (PI. 3, Figs 1, 2;
specimen length 85 /urn)

Sierre Leone Kitton, BM 1890, lectotype, BM 1892; Sierre Leone, BM 2235.
Although there is a drawing of this species in the Greville manuscript collection, there is no
indication which slide or specimen was used for the original description. There are three relevant
slides in the collection: BM 1890 (specimen no. 2), BM 1892 (one specimen at MF 30/19), and
BM 2235 (two specimens at MF 29/31 and MF 21/15). BM 1890, specimen no. 2, is designated as
lectotype. There is an additional drawing in the manuscript collection of an apparently
unpublished figure from BM 1892 (MF 30/19).

5. A. delicatula Grev. in Edinb. Newphil. J. 18: 39, figs 15, 16 (1863); in Trans, hot. Soc. Edinb.
7: 539, figs 15, 16 (1863). (PI. 3, Figs 3-5; specimen length 57-5 /am)

Woodlark Isld. Roberts 1863, BM 2717, BM 2719, syntypes.

There are two illustrations given with the published description, both of which are in the Greville
manuscript collection. The valve view corresponds to BM 2717 (MF 25/24; not illustrated) and
the girdle view corresponds to BM 2719 (MF 16/32; illustrated). These are syntypes.

6. A. eximia Grev. in Edinb. Newphil. J. 18: 38, fig. 13 (1863); in Trans, hot. Soc. Edinb. 7: 538,
fig. 13 (1863). (PI. 3, Fig. 7; specimen length 132-5 /xm)

Curteis Straits Queensland Dr Roberts [18]63, BM 2626, holotype.

7. A. hyalina Grev. in Ann. Mag. nat. Hist. Ill, 16: 6, fig. 11 (1865). (PI. 2, Fig. 3; specimen
length 45 -5 /xm)

Hong Kong 1864 no. 3, BM 4507, syntypes.

No specimen is indicated as present on this slide by Greville, although the drawing in the
Greville manuscript collection identifies this slide as the one he used. The poor quality of the
micrograph is due to the dry mounted slide and a very thick coverslip. The specimens on this
slide should be considered syntypes.

8. A. jolisiana Grev. in Edinb. Newphil. J. 18: 186, fig. 11 (1863); in Trans, hot. Soc. Edinb. 7:
579, fig. 11 (1863). (PI. 2, Fig. 7; specimen length 67-5 ^m)

Curteis Straits Dr Roberts 1862, BM 2629, holotype.

Two specimens are indicated on the slide; the drawing in the Greville manuscript collection
indicates that MF 13/22 is the holotype, whilst MF 15/24 is an isotype. The micrograph presented
here is of the holotype, MF 13/22.

9. A. kuetzingiana Grev. in Edinb. Newphil. J. 18: 184, fig. 6 (1863); in Trans, hot. Soc. Edinb.
7: 577, fig. 6 (1863). (PI. 4, Figs 1, 2; specimen length 110 ,um)

Curteis Straits Queensland Dr Roberts 1862, BM 2630, holotype.

10. A. lata Grev. in Edinb. Newphil. J. 18: 38, fig. 14 (1863); in Trans, hot. Soc. Edinb. 7: 539,
fig. 14 (1863). (PI. 4, Figs 3, 4; specimen length 102-5 /xm)

Curteis Straits Dr Roberts 1863, BM 2615, holotype.

11. A. lepida Grev. in Ann. Mag. nat. Hist. Ill, 16: 5, fig. 10 (1865). (PI. 3, Figs 8, 9; specimen
length 145 /xm)

Hong Kong, BM 4520, holotype.

The drawing in the Greville manuscript collection gives a MF location and a slide reference

number. The slide reference number is Greville's annotation referring to his working slides

and is marked simply as no. 1. Slide BM 4520 is labelled T and, in pencil, 'Hong Kong'.

The specimen located at MF 22/20 is identical to the published figure and is therefore the

holotype.

12. A. lineata Grev. in Edinb. Newphil. J. 18: 40, fig. 19 (1863); in Trans, hot. Soc. Edinb. 7:
540, fig. 19 (1863). (PI. 4, Figs 5, 6; specimen length 82-5 /*m)

Woodlark Isld. Roberts, 1863, BM 2717, holotype.

This species has been studied in detail by Paddock & Sims (1980: 169, esp. 171) and is the type of

their new genus Undatella Paddock & P. A. Sims.

6 D. M. WILLIAMS

13. A. meneghiniana Grev. in Edinb. New phil. J. 18: 184, fig. 7 (1863); in Trans, hot. Soc.
Edinb. 7: 577, fig. 7 (1863). (PI. 4, Figs 7, 8; specimen length 97-5 /im)

Curteis Strait, BM 2888, holotype.

Although there is only one illustration on the Greville manuscript drawing, two slide numbers
are given. BM 2601 (MF 21/21) locates only a fragment of a valve. Additionally, the annotation
is followed by an exclamation mark. BM 2888 (MF 34/24) corresponds exactly with that
illustrated and it is concluded that this is the holotype.

14. A. nitida Grev. in Edinb. New phil. J. 18: 40, fig. 18 (1863); in Trans, hot. Soc. Edinb. 7: 540,
fig. 18 (1863). (PI. 5, Figs 1, 2; specimen length 77-5 /am)

Curteis Straits Queensland, BM 2614, holotype.

Two specimens are indicated on the slide; from comparison of each with the published drawings,

MF 20/13 is the holotype, whilst 28/23 is a paratype.

15. A. oblonga Grev. in Trans, microsc. Soc. Lond. 11: 20, fig. 15 (1863). (PI. 3, Fig. 6; Detail
PI. 5, Figs 3-5; specimen length 150 /am)

Harvey Bay, Dr Roberts 1862, BM 2671, lectotype, BM 2677; Harvey Bay, Queensland, BM
2668, BM 2676, BM 2679; Harvey Bay, Queensland, Dr Roberts 1862, BM 2675.
The drawing from the Greville manuscript collection is missing and, from the annotations on the
slides, it is difficult to establish which specimen Greville used for his original illustration. BM
2671 (MF 24/12) has been selected as a lectotype. Other specimens are: BM 2671 (MF 20/17),
BM 2668 (MF 21/29 and 16/23), BM 2675 (MF 29/27), BM 2676 (MF 25/26), BM 2677 (MF 22/18
and 33/19), and BM 2679 (MF 35/22). Interestingly, Greville states in the protologue that a
girdle view was difficult to find. BM 2677 (MF 33/19) is identified by Greville and is a girdle view.

16. A? paradoxa Grev. in Edinb. New phil. J. 18: 41, fig. 21 (1863); in Trans, hot. Soc. Edinb. 7:

541, fig. 21 (1863). (PI. 5, Fig. 7; specimen length 80 /urn)
Curteis Straits Queensland, BM 2616, holotype.

17. A? rabenhorstiana Grev. in Edinb. New phil. J. 18: 185, fig. 12 (1863); in Trans, hot. Soc.
Edinb. 7: 578, fig. 12 (1863). (PI. 5, Fig. 8; specimen length 120 /am)

Curteis Straits Dr Roberts 1862, BM 2624, holotype.

18. A. rectangularis Grev. in Edinb. New phil. J. 18: 185, fig. 10 (1863); in Trans, hot. Soc.
Edinb. 7: 578, fig. 10 (1863). (PI. 5, Fig. 6; specimen length 72-5 /am)

Woodlark Island Dr Roberts 1862, BM 2715, holotype.

19. A? superba Grev. in Edinb. New phil. J. 18: 39, fig. 17 (1863); in Trans, hot. Soc. Edinb. 7:
539, fig. 17 (1863). (PI. 6, Figs 1-3; specimen length 232-5 /am)

New Caledonia, BM 2641, holotype.

20. A. thwaitesiana Grev. in Edinb. New phil. J. 18: 183, fig. 9 (1863); in Trans, hot. Soc. Edinb.
7: 576, fig. 9 (1863). (PI. 5, Figs 9, 10; specimen length 72-5 /am)

Curteis Straits Dr Roberts 1862, BM 2603, holotype.

21. A? undulata Grev. in Edinb. New phil. J. 18: 42, fig. 22 (1863); in Trans, hot. Soc. Edinb. 7:

542, fig. 22 (1863). (Not illustrated)

Curteis Straits Queensland Dr Roberts 1862, BM 2611, holotype.

22. A. venusta Grev. in Ann. Mag. nat. Hist. Ill, 16: 6, fig. 12 (1865).
Hong Kong/. L. Palmer, BM 4505, holotype.

The holotype, based on data given on the drawing in the Greville manuscript collection,
corresponds to a broken slide. The coverslip has detached from the glass slide and although this
can be re-mounted, it is impossible to re-trace the MF numbered specimen once this has been
done.

V. AMPHITETRAS Ehrenb.

The relationships of Amphitetras Ehrenb., Amphipentas Ehrenb., and Triceratium Ehrenb.,
within the Eupodiscaceae have been discussed by Ross & Sims (1970), but no generic limits were

GREVILLE DIATOM TYPES 7

defined. Simonsen (1968) implies in his index to the Greville papers that both genera belong in
Triceratium.

1. A. elegans Grev. in Trans, microsc. Soc. Lond. 14: 9, fig. 24 (1866). (PI. 6, Figs 4, 8; specimen
length 72-5 /zm)

Monterey L. H. no. 31, BM 10007, holotype.

2. A. minuta Grev. in Trans, microsc. Soc. Lond. 9. 77, fig. 11 (1861). (PI. 6, Fig. 5; specimen
length 40 /xm)

Nottingham deposit U.S., BM 2187, holotype.

3. A. nobilis Grev. in Trans, microsc. Soc. Lond. 13: 105, fig. 27 (1865).
Red Sea L. H. no. 34, BM 10009.

This slide has suffered considerable damage where the coverslip has detached from the glass
slide. An attempt to re-mount the coverslip has not been successful.

4. A. parvula Grev. in Edinb. New phil. J. 18: 37, fig. 12 (1863); in Trans, hot. Soc. Edinb. 7:
537, fig. 12 (1863). (PI. 6, Figs 6, 7; specimen diameter 22-5 /xm)

New Caledonia Dr Roberts 1863, BM 2664, holotype.

This is a specimen of Perissonoe cruciata (Janisch & Rabenh.) G. W. Andrews & Stoelzel
(Andrews & Stoelzel, 1984). Greville mistook the air-bubble in the centre of the specimen as
part the valve structure. A search on the rest of the slide revealed several good specimens of
P. cruciata, of which one is located at England Finder 36K2 [rear].
The correct name for this species is given in the appendix.

5. A. producta Grev. in Trans, microsc. Soc. Lond. 10: 94, fig. 11 (1862). (PI. 6, Fig. 9; specimen
diameter 52-5 /xm)

Manilla shell scrapings G. M. Browne, BM 2382, lectotype.

There is no drawing of this species in the Greville manuscript collection. In the protologue,
Greville gives two localities: 'Manilla . . . George Mansfield Browne, Esq. Dredged off St.
Helena; Dr. Wallich.' Both these localities are represented in the Greville slide collection. BM
2382, from Manilla, has a few selected specimens of which one is A. producta. The two other
slides, BM 2579 and BM 3228, do not have specimens of A producta indicated on the label nor
were any located on the slides. BM 2382 should therefore be considered as the lectotype.

6. A. punctata Grev. in Trans, microsc. Soc. Lond. 10: 29, fig. 8 (1862). (PI. 6, Fig. 10; specimen
diameter 55 /xm)

Ceylon Dr Macrae, BM 2391, holotype.

7. A. radiata Grev. in Trans, microsc. Soc. Lond. 10: 28, fig. 7 (1862). (PI. 7, Figs 1,2; specimen
diameter 147-5 /am)

Peru Guano Dr Macrae, BM 2399, holotype.

VI. AMPHORA Ehrenb. ex Kiitz.

Many species of Amphora are currently being revised by Schoeman & Archibald (1986a-c,
1987fl, b} and Schoeman, Archibald & Sims (1986). Additionally, one of Greville's species has
been placed in the new genus Undatella Paddock & P. A. Sims.

1. A. flexuosa Grev. in Edinb. New phil. J. 18: 183, fig. 4 (1863); in Trans, hot. Soc. Edinb. 7:
576, fig. 4 (1863). (PI. 7, Fig. 5; specimen length 77-5 /*m)

Curteis Straits Queensland Dr Roberts [18]63, BM 2626, holotype.

2. A. magnified Grev. in Edinb. New phil. J. 18: 182, fig. 1 (1863); in Trans, hot. Soc. Edinb. 7:
575, fig. 1 (1863). (PI. 7, Figs 8, 9; specimen length 117-5 /xm)

Curteis Straits Dr Roberts, 1863, BM 2713, holotype, BM 2593 (figured in Paddock & Sims
(1980) as pi. 6, figs 48, 50), BM 2600, BM 2602, BM 2613, BM 2616, BM 2626, BM 2632, BM
2635, BM 2638, BM 2639, BM 2881 (figured in Paddock & Sims (1980) as pi. 6, figs 47, 49), BM
2882, BM 2885, BM 2886, BM 2888, BM 4644, BM 4659.

8 D. M. WILLIAMS

Greville indicated on the drawing in the Greville manuscript collection the exact slide he
referred to for the illustration of this species. The slide is not annotated. There are a
considerable number of specimens in the collection; all should be considered as paratypes. It is
probable that Greville drew upon this large number of specimens for the description. For a
recent detailed treatment of this species see Paddock & Sims (1980).

3. A. pulchra Grev. in Edinb. New phil. J. 18: 182, fig. 2 (1863); in Trans, hot. Soc. Edinb. 7:
575, fig. 2 (1863). (PI. 7, Figs 6, 7; specimen length 107-5 /am)

Curteis Straits 1863, BM 2633, holotype; BM 2597, BM 2636, BM 2884, BM 2889, BM 4665.
The drawing in the Greville manuscript collection indicates BM 2633 (MF 24/17) as the
holotype. For a recent detailed treatment of this species see Paddock & Sims (1980). They
include micrographs of the holotype (pi. 2, fig. 11), along with scanning electron micrographs of
some Recent material. See also Amphiprora lineata.

4. A. sarniensis Grev. in Trans, microsc. Soc. Lond. 10: 95, fig. 12 (1862). (Not illustrated)
Guernsey Dr Wallich, BM 2530, BM 2531 lectotype, BM 2532.

There is no drawing in the Greville manuscript collection of this species and three slides are
possible sources of the holotype. Schoeman & Archibald (19876) have studied the genus
Amphora extensively, including this species. They have designated as lectotype BM 2531.

5. A. sinuata Grev. in Trans, hot. Soc. Edinb. 7: 576, fig. 5 (1863); in Edinb. New phil. J. 18:
183, fig. 5 (1863). (PI. 7, Fig. 4; specimen length 50 /xm)

Curteis Straits Queensland Dr Roberts 1862, BM 2595, holotype.

6. A. undulata Grev. in Edinb. New phil. J. 18: 182, fig. 3 (1863); in Trans, hot. Soc. Edinb. 7:
575, fig. 3 (1863). (PI. 7, Fig. 3; specimen length 65 /xm)

Curteis Straits Queensland, BM 2616 (MF 27/36), holotype; BM 2597, BM 2631.

VII. ARACHNOIDISCUS Ehrenb .

1. A. grevilleanus Hardman ex Grev. in Trans, microsc. Soc. Lond. 13: 47, fig. 7 (1865). (Not

illustrated)

Cambridge Estate Barbadoes Johnson 1864, BM 3278; Barbadoes 1865, BM 3284; Barbadoes

Johnson, BM 3445; Barbadoes Johnson, BM 3463; Springfield Estate Barbadoes L. Hardman

1864, BM 3343.

No indication is given on the drawing as to which slide was used for the original description.

Several localities were given in the protologue, all of which are represented by slides. All

specimens are obscured by debris, with most either fragmented or obscured by some other

material.

VIII. ASTERIONELLA Hassall

1. A. synedraeformis Grev. in Ann. Mag. nat. Hist. Ill, 16: 4, figs 5, 6 (1865). (Not illustrated)
Hong Kong Harbour/. L. Palmer 1863, BM 3229; Hong Kong Harbour T. G. R. [ylands], BM
4438, syntypes.

IX. ASTEROLAMPRA Ehrenb.

The most recent revision of the genus by Gombos (1980) has intepreted a few species in relation
to their phylogenetic position in the family Asterolampraceae. After making a detailed study of
the 'rays' by SEM, Gombos drew attention to Greville's descriptive terminology. Where
Greville had referred to 'umbilical lines' as the side walls of the 'rays', Simonsen (in Gombos,
1980) suggested that they be replaced with the term 'separating line' , and umbilicus should be
replaced by junction point. The quotes below from Greville's descriptions have been made
verbatim. Nevertheless, the above modern terminology should be borne in mind.

Many of Greville's species would be placed in the 'genus' Liostephania Ehrenb., which is
apparently preserved casts of Asterolampra valves. A review by Payne (1922) and a recent short

GREVILLE DIATOM TYPES 9

paper by Hanna & Brigger (1970) show a number of Greville's species as such casts: A. laevis, A.
aemulans, A. simulans, A. pulchra, A. kittoniana, A. stellulata, A. dubia, A. ambigua, A. aliena,
and A. scutula.

1. A. aemulans Grev. in Trans, microsc. Soc. Lond. 10: 52, figs 34, 35 (1862). (PI. 8, Figs 1, 2;
specimen diameter 45 Aim)

Barbadoes deposit, BM 2031, lectotype (MF 16/26), BM 2030.

There is no drawing in the Greville manuscript collection. Two slides are indicated by Greville
for this species: BM 2031 and BM 2030. BM 2030 has only a fragmented valve, whilst BM 2031
has two entire valves, MF 16/26 and 34/28. The former has been photographed as the latter is
partially obscured by another diatom.

2. A. affinis Grev. in Trans, microsc. Soc. Lond. 10: 45, figs 7-9 (1862). (PI. 8, Figs 3-6)
Barbadoes deposit, BM 2015, BM 2030, BM 2036, BM 2063, BM 2068, BM 2069.

There are no drawings in the Greville manuscript collection. In the protologue Greville refers to
several valves he inspected. It is not clear which of these specimens were used for the series of
illustrations. Some of the valves observed on the slides can be matched with those of Greville's
descriptions: 'I have only seen one individual in which umbilical cellules were entirely absent
[BM 2030 (24/25) Greville's fig. 7; PI. 8, Fig. 5; specimen diameter 45 /am]. The valve, fig. 9, is
the largest disc belonging to the genus which has occurred in the deposit [BM 2015 (30/10) (PI. 8,
Fig. 5; specimen diameter 82-5 Aim)]'. Greville's figure 8 is probably BM 2015 (MF 19/25; PI. 8,
Fig. 6; specimen diameter 60-5 joon).

Additionally, the label of BM 2068 is annotated by Greville as having two specimens, but none
were found corresponding to the MF numbers. The coverslip may have moved and the
specimens no longer correspond to their original position.

3. A? aliena Grev. in Trans, microsc. Soc. Lond. 10: 55, fig. 46 (1862). (PI. 9, Fig. 1; specimen
diameter 50 AUTI)

Barbadoes deposit, BM 2032, holotype?

There is no drawing in the Greville manuscript collection. Two slides are indexed by Greville as
having specimens of A. aliena. BM 2604 has a note attached to the label which states: The
Asterolampra is allied to A. aliena but differs somewhat.' Therefore, the other slide, BM 2032, is
probably the holotype. Another specimen exists in the Hardman collection, BM 10051.

4. A. ambigua var. a Grev. in Trans, microsc. Soc. Lond. 10: 54, figs 42-44 (1862).
Barbadoes deposit, BM 2068, holotype?

In the protologue Greville states: 'I am indebted to my friend Mr. Rylands for the use of two
drawings, from which I have made the figs. 42 and 44'. Neither of these figures are extant in the
Greville manuscript collection or in any of Ryland's papers so far examined. The specimen on
BM 2068 could not be located.

5. A. ambigua var. b Grev. in Trans, microsc. Soc. Lond. 10: 54, fig. 45 (1862). (PI. 9, Fig. 2;
specimen diameter 40 //,m)

Barbadoes deposit, BM 2064 (MF 17/19), holotype?

This is the only slide in the Greville collection referable to var. b and is probably the holotype.

6. A. brebissoniana Grev. in Trans, microsc. Soc. Lond. 8: 114, fig. 9 (1860).
Monterey Stone Arnott (647), BM 1941; Monterey Stone Arnott (6472), BM 1942.
Neither of these slides corresponds to specimens of the above species.

7. A. concinna Grev. in Trans, microsc. Soc. Lond. 10: 46, figs 10-12 (1862). (PI. 9, Figs 5, 7;
specimen diameter 70 Aim)

Barbadoes deposit, BM 2017, lectotype (MF 23/37), BM 2016, BM 2063.
Greville notes that 'I have only seen four specimens, all with seven rays, and destitute of
umiblical cellules.' These four specimens are: BM 2016 (MF 22/21), BM 2017 (MF 23/37), and
BM 2063 (MF 35/31 and MF 16/32). The micrograph (BM 2017) is from the best of these four.
The other specimens are obscured by either diatoms or debris.

10 D. M. WILLIAMS

8. A. crenata Grev. in Trans, microsc. Soc. Lond. 10: 47, figs 14-16 (1862). (PI. 9, Fig. 6;
specimen diameter 47-5 /xm)

Barbadoes deposit, BM 2048, lectotype, BM 2022, BM 2024, BM 2033, BM 2064, BM 2066.
Several specimens are indicated in the Greville slide collection. BM 2048 has been chosen as
lectotype because it presents a whole specimen.

9. A. dallasiana Grev. in Trans, microsc. Soc. Lond. 8: 115, fig. 10 (1860).
Bermuda Tripoli, BM 1287, BM 1728; Nottingham dept. U.S. , BM 2196.
No specimens could be located on the above slides.

10. A. decora Grev. in Trans, microsc. Soc. Lond. 10: 45, figs 4-6 (1862). (PI. 10, Figs 1-4)
Barbadoes deposit, BM 2015, BM 2021 , BM 2024, BM 2031, BM 2037, BM 2038, BM 2064, BM
2065, BM 2067, BM 2068, BM 2170, BM 2073.

According to Greville, this species exhibits considerable variation in pattern. He illustrates this
in his text by stating: 'I have examined a large suite of specimens, and in twenty-five, of which I
have taken notes, the number [of rays] varied from five to fourteen, and that of umbilical cellules
is equally uncertain.' Not all of the 25 specimens that Greville refers to have been located in his
collection. However, a large number have been discovered which are usually annotated on the
slide label with the number of rays that are present, i.e. BM 2015, 6 rays, BM 2021, 14 rays, BM
2024, 6 rays, BM 2031, 8 rays, BM 2038, 10 rays, BM 2067, two specimens with 10 rays and 6
rays. Finally, BM 2037 has no annotation but has 6 rays, BM 2064 has three specimens, and BM
2065 has 10 rays. BM 2068 is annotated as having no cellules.

It is possible to compare the published figures with the specimens. Greville's figure 4 is
probably BM 2024, but it is difficult to be certain as the specimen lies at an angle. This may
account for the overestimation of the number of central cellules in this specimen. Greville's
figure 5 is BM 2015 (PI. 10, Figs 2-4; specimen diameter 55 /xm) and Greville's figure 6 is BM
2021 (PI. 10, Fig. 1; specimen diameter 45 /xm).

11. A. decorata Grev. in Trans, microsc. Soc. Lond. 10: 46, fig. 13 (1862). (PI. 10, Figs 5-8;
specimen diameter 72-5 /xm)

Barbadoes deposit, BM 2018, holotype?

In the protologue Greville states that: 'Only one valve of this exquisitely beautiful disc has been

found.' As BM 2018 is the only slide with this taxon, it is probably the holotype.

12. A. dubia Grev. in Trans, microsc. Soc. Lond. 10: 54, fig. 41 (1862).
Barbadoes deposit, BM 2037, holotype?

There is no drawing of this specimen in the Greville manuscript collection and only one slide
number is given in the catalogue index. I have been unable to locate this specimen. It remains to
be seen if this slide is the holotype.

13. A. eximia Grev. in Trans, microsc. Soc. Lond. 13: 99, fig. 10 (1865). (PI. 10, Figs 9, 10;
specimen diameter 70 /xm)

Barbadoes, L.H. no. 1139, BM 10895, holotype?

Greville indicates that he obtained his specimen from Hardman. It is possible that this slide is the

holotype.

14. A. hiltoniana Grev. in Trans, microsc. Soc. Lond. 8: 117, fig. 15 (1860). (PI. 11, Figs 1, 2;
specimen diameter 90 /xm)

Ind. Oc. Soundings Captain Pullen 2200 fathom, BM 1927, BM 1880, BM 1882, BM 1923, BM
1925, BM 1926, BM 1937, BM 1951; Algoa Bay Guano, BM 1678, syntypes.
Greville gives two localities in the protologue for this species. However, in the text he notes 'In
the Indian one, which I have figured with nineteen rays' (see his figure 15), indicating which
gathering was used for the drawing. There is no specimen identified as A. hiltoniana on the
Algoa Bay slides. Greville refers to these specimens in an earlier account (Greville, 1859b: 160).
BM 1678 has a single specimen of an Asterolampra species (probably A. marylandica Ehrenb.).
This has the simple lines that Greville referred to and may be the Algoa Bay specimen he used in
the description of A. hiltoniana. The micrograph is from BM 1880.

GREVILLE DIATOM TYPES 11

Rattray (1888a) lists the African locality in his description of this species, which would suggest
he had examined Greville's material, but there are no annotations to the Greville slides as was
Rattray 's usual practice.

15. A. kittoniana Grev. in Trans, microsc. Soc. Lond. 10: 53, fig. 39 (1862). (PI. 11, Fig. 3;
specimen diameter 40 /am)

Barbadoes deposit, BM 57803, holotype; BM 2069.

Greville noted in the protologue: 'For a drawing of this elegant little species I am indebted to my
very obliging correspondent, Mr. Kitton, of Norwich, who alone has discovered it.' In the
Wynne Baxter collection there is a series of slides that were purchased from the executors of
Kitton's estate. Among these slides, BM 57803 (Wynne Baxter 3181) is from Barbados. In
volume 4 of the catalogue that accompanied Wynne Baxter's slide collection, there is an
annotation against this slide number: 'This is the original specimen from which Greville took his
description and figure.' As this slide was originally in Kitton's herbarium it is safe to assume this
is the holotype (MF 37/28).

16. A. laevis Grev. in Trans, microsc. Soc. Lond. 10: 51, fig. 33 (1862). (PI. 11, Figs 4, 5;
specimen diameter 27-5 /am)

[Barbadoes deposit], BM 2072, holotype?

There is no drawing of this specimen in the Greville manuscript collection. BM 2072 is the only

slide in Greville's collection with this name on the label.

17. A. moronensis Grev. in Q. Jl microsc. Sci. II, 3: 230, fig. 8 (1863).
Moron nr. Seville Norman 1862, BM 2698, holotype?

There is a drawing in the Greville collection which indicates that specimen MF 18/30 of slide BM
2698 is the one used for the illustration. On the manuscript drawing the number 33/16 appears in
the top corner but it has been crossed out. Neither MF numbers locate specimens on this slide.
The holotype has yet to be discovered.

18. A. pulchra Grev. in Trans, microsc. Soc. Lond. 10: 53, figs 37, 38 (1862). (PI. 11, Figs 6, 7;
specimen diameter 57-5 /mi)

Barbadoes deposit, BM 2036, lectotype, BM 2062.

Greville gave two illustrations for this species corresponding with the above two slides. BM 2036
is the specimen in figure 38; BM 2062 (MF 13/22) is the specimen in figure 37. BM 2036 has been
chosen as lectotype as it is an entire valve.

19. A. punctata Grev. in Trans, microsc. Soc. Lond. 10: 51, fig. 32 (1862). (PI. 11, Figs 8, 9;
specimen diameter 70 /am)

Barbadoes deposit, BM 2252, holotype?; [Barbadoes deposit], BM 2064; Barbadoes deposit,
BM 2072, BM 2086.

All the annotated slides in Greville's collection have specimens with only six rays. BM 2252 has
seven rays, as does the published illustration. This probably indicates that it is the holotype.

20. A. ralfsiana Grev. in Trans, microsc. Soc. Lond. 10: 50, fig. 31 (1862). (PI. 12, Fig. 1;
specimen diameter 65 /am)

Barbadoes deposit, BM 2039, lectotype, BM 2015, BM 2063, BM 2065, BM 2070, BM 2072, BM
2075, BM 2079, BM 2109, BM 2200.

All the slides from the type locality are of rather poor quality. BM 2039 was chosen as lectotype
as it is the closest to that represented in the published figure.

21. A. roperiana Grev. in Trans, microsc. Soc. Lond. 8: 120, fig. 14 (1860). (PI. 12, Fig. 2;
specimen diameter 85 /am)

Ind. Oc. Soundings Captain Fallen 2200 fathom, BM 1924 lectotype, BM 1874, BM 1882, BM
1930, BM 1931, BM 1933, BM 1934, BM 1951, BM 4417.

Greville states in the protologue that he made 'examination^] of above a dozen specimens' and
'the individual figured is the most perfect but by no means the largest in my cabinet.' The only
specimen that appears to be complete is that on BM 1924 (MF 30/17) and hence is chosen as the
lectotype. BM 1930 is annotated 'large fragment' and BM 1933 is annotated '.0066'.

12 D. M. WILLIAMS

22. A. rotula Grev. in Trans, microsc. Soc. Lond. 8: 111, fig. 5 (1860).

Monterey Stone Arnott (647), BM 1941; Monterey Stone Arnott (6472), BM 1942, BM 1967.
Greville made a number of drawings of this species but there is no indication which slide was
used for the published illustration. BM 1941 has been chosen as lectotype.

23. A. rylandsiana Grev. in Trans, microsc. Soc. Lond. 10: 49, figs 28, 29 (1862). (PI. 12, Fig. 3;
specimen diameter 40 /xm)

Barbadoes deposit, BM 2021, BM 2023, BM 2025, BM 2063, BM 2064, BM 2065, BM 2068.
The two figures Greville published may correspond to the following slides: figure 28 possibly to
BM 2021 (PI. 12, Fig. 3) and figure 29 possibly to BM 2068 (not illustrated).

24. A. scutula Grev. in Trans, microsc. Soc. Lond. 10: 52, fig. 47 (1862). (PI. 12, Fig. 4;
specimen diameter 35 /Am)

Barbadoes deposit Johnson 1861, BM 2473, holotype.

This is the only slide of this species in the Greville collection and is probably the holotype. The

slide is rather thick, allowing only low magnification and rather poor focus.

25. A. shadboltiana Grev. in Trans, microsc. Soc. Lond. 8: 121, fig. 19 (1860). (PI. 12, Fig. 5;
specimen diameter 62-5 /am)

Ind. Oc. Soundings Captain Fallen 2200 fathom, BM 1935 lectotype, BM 1936.
In the protologue Greville states: 'Mr. Roper very kindly sent both his accurate drawing and the
slide containing the valve of this interesting species for my inspection.' It is not clear from the
slides which, if any, came from Roper. As Greville states that the specimens he discovered from
his own collection were identical to that of Roper's, BM 1935 has been chosen as lectotype. BM
1936 has the label annotated with A. shadboltiana. However, the name is followed by a query
and suggests doubt in Greville's mind. There are a number of slides in the Roper collection (BM)
of this species (BM 21659, BM 21671, BM 21667).

26. A. simulans Grev. in Trans, microsc. Soc. Lond. 10: 52, fig. 36 (1862). (PI. 12, Fig. 6;
specimen diameter 57-5 ^tm)

Barbadoes deposit, BM 2021, holotype?

There is no drawing of this specimen in the Greville manuscript collection. The above slide is the

only one in Greville's collection annotated for this species.

27. A. stellata Grev. in Trans, microsc. Soc. Lond. 8: 124, fig. 20 (1860). (Not illustrated)
Indian Ocean soundings, BM 1938, BM 1939, BM 1951, syntypes.

Greville only suggested a provisional name for this taxon. However, it has been accepted by
many subsequent authors.

28. A. stellulata Grev. in Trans, microsc. Soc. Lond. 10: 54, fig. 40 (1862). (PI. 12, Fig. 7;
specimen diameter 32-5 /xm)

Barbadoes deposit, BM 2016, holotype?, BM 2032.

Greville states in the protologue that he has 'only seen two examples [of this species] one of
which has seven, the other nine rays.' On each of the slides above there is one specimen; both
should be considered syntypes. However, the illustration is of a seven-rayed specimen which is
present on BM 2016 and this should probably be considered the holotype.

29. A. variabilis Grev. in Trans, microsc. Soc. Lond. 8: 111, figs 6-8 (1860). (PI. 9, Figs 3, 4; PI.
12, Figs 8-12; PI. 13, Figs 1, 2; specimen diameter 45-60 //-m)

Monterey Walker- Arnott, BM 1941, BM 1942, BM 1971, BM 2008, BM 2012, BM 2250, BM

2253, BM 2255.

In the protologue Greville gives an extended discussion of the variability of this species, giving

six types of variation pattern. Of these, three are illustrated and the type of variation in pattern

stated:

(1) Seven rays, one line is simple, six united in triplets, only three radiate from the central
point.

(2) Eight rays.

GREVILLE DIATOM TYPES 13

(3) Eight rays , one simple line , two forked , one triplet , four lines radiating from the centre , but
by a different combination of rays.

(4) Nine rays, two simple lines, two forked and one triplet (Greville's fig. 7).

(5) Ten rays, no single line but two forked and two triplets.

(6) Eleven rays, one simple line, two triplets and one quadruplet (Greville's fig. 8).

As this species involves such a diverse series of forms, it was decided not to propose a lectotype
until the series has been studied thoroughly.

30. A. vulgaris var. a Grev. in Trans, microsc. Soc. Lond. 10: 47, figs 17-20 (1862). (PI. 13, Figs
3-9; specimen diameter 55-137 /xm)

Barbadoes deposit, BM 2017, BM 2018, BM 2020, BM 2023, BM 2024, BM 2035, BM 2045?,

BM 2049, BM 2063, BM 2064.

Greville illustrated this taxon with a wide range of variation.

31. A. vulgaris var. b Grev. in Trans, microsc. Soc. Lond. 10: 47, fig. 21 (1862). (PI. 13, Fig. 10;
specimen diameter 52-5 /xm)

Barbadoes deposit, BM 2022, holotype.

32. A. vulgaris var. c Grev. in Trans, microsc. Soc. Lond. 10: 47, fig. 22 (1862).
Barbadoes deposit, BM 2020.

This is the only slide annotated for this variety and is probably the holotype. Unfortunately no
specimen could be located.

33. A. vulgaris var. d Grev. in Trans, microsc. Soc. Lond. 10: 47, figs 23, 24 (1862). (PI. 13, Figs
11, 12; specimen diameter 57-5 /xm)

Barbadoes deposit, BM 2015, lectotype; [Barbadoes], BM 2026, BM 2030, BM 2035, BM 2063,
BM 2070.

34. A. vulgaris var. e Grev. in Trans, microsc. Soc. Lond. 10: 47, fig. 25 (1862). (= A. splendida
Grev. ex Rattray in Trans. R. microsc. Soc. 1890: 648 (1890).) (PI. 14, Fig. 6; specimen diameter
40 /xm)

Barbadoes deposit, BM 2094, holotype.

Greville originally listed this species as A. vulgaris var. e, with a note in the text saying 'the most
extreme variety is one which I first considered distinct, and named A. splendida (fig. 25) and
even now I am not quite satisfied about its true position.'

35. A. wallichiana Grev. in Trans, microsc. Soc. Lond. 8: 115, fig. 11 (1860). (PI. 14, Fig. 5;
specimen diameter 77-5 /xm)

Bermuda Tripoli Bailey 1856, BM 1330.

The protologue has E. W. Dallas as collector, but the slide indicates J. W. Bailey. There is no
drawing in the Greville manuscript collection and hence it is doubtful that this is Greville's
original material.

X. ASTEROMPHALUSEhrenb.

1. A. elegans Grev. in Q. Jl microsc. Sci. 7: 161, fig. 6 (1859). (PI. 14, Fig. 4; specimen diameter

72 /xm)

Calif orn. guano Norman 8.54, BM 1778, holotype.

XI. A ULACODISCUS Ehrenb .

A detailed monograph of this genus has been undertaken by Burke & Woodward (1963-1970)
and continued by Burke (1970-1974). This revision is based on light microscope observations. A
recent SEM study on Aulacodiscus was presented by Ross & Sims (1970). One of the species
they examined was A. grevilleanus .

1. A. amoenus Grev. in Trans, microsc. Soc. Lond. 12: 10, fig. 3 (1864). (PI. 14, Fig. 3; specimen
diameter 104-5 /xm)

14 D. M. WILLIAMS

Cambridge Estate Barbadoes Johnson, BM 2860, holotype; BM 2862, 2863, 3070, 3071, 3122,

3204,3205.

There are several specimens in the slide collection but only one is labelled as holotype. BM 2860

(MF 26/25) is the holotype. The mountant has deteriorated considerably preventing the use of

high magnification. BM 3204 and BM 3205 are syntypes of A. amoenus var. subdecorus Rattray .

2. A. angulatus Grev. in Trans, microsc. Soc. Lond. 11: 71, fig. 15 (1863). (PI. 14, Fig. 1;
specimen diameter 105 /xm)

Cambridge Barbadoes Johnson 1863, BM 2744, holotype, BM 2895; Barbadoes Johnson 1863,
BM 2941; Cambridge Barbadoes Johnson 1863, BM 2981.

There is a series of manuscript notes which indicate that Greville used BM 2744 as the holotype.
The specimen is at an angle and is difficult to photograph clearly. A good specimen has been
photographed from BM 2981 (PI. 14, Fig. 2; specimen diameter 112-5 /xm).

3. A. attenuatus Rattray inJlR. microsc. Soc. 1888: 356, pi. 5, fig. 2 (1888). (A. elegans Grev.
nom. herb., non A elegans Grove & Sturt (1887).) (PI. 15, Figs 10, 11; specimen diameter 97-5

Cambridge Barbadoes Johnson June 63, BM 2897, lectotype; Cambridge Barbadoes Johnson

1862, BM 2758; Cambridge Barbadoes Johnson 1864, BM 3203; Barbadoes Johnson 1865, BM

3449.

Rattray described this species as new. However, the specimens annotated in the Greville

herbarium are all marked as A. elegans by Greville. Rattray, who was aware of preoccupation,

proposed a new name.

There is no indication which of these slides Rattray used for his illustration (which is a portion
of a valve). BM 2897 has been chosen as the lectotype.

4. A. cinctus Grev. ex Rattray in JIR. microsc. Soc. 1888: 365 (1888). (PI. 15, Figs 6, 9; specimen
diameter 100 /xm)

Barbadoes 1864, BM 3031, lectotype; Barbadoes Johnson 1863, BM 2934; Cambridge Estate
Barbadoes Johnson 1863, BM 2979; Cambridge Estate Barbadoes Johnson 1864, BM 3069, BM
3170, BM 3249, BM 3250; Barbadoes 186$, BM 3245; Barbadoes Johnson 1864, BM 3268, BM
3269, BM 3283; Cambridge Barbadoes, BM 4779, BM 4784.

Although Greville named this species he did not describe or publish it. Rattray (18886), in his
revision of Aulacodiscus , described the species for the first time with acknowledgement to
Greville. There is no indication in Greville's notes or in Rattray's text which specimen was used
for the type description and so all the above, annotated by Greville, can be considered as
syntypes. BM 3031 has been selected as the lectotype.

5. A. decorus Grev. in Trans, microsc. Soc. Lond. 12: 82, fig. 2 (1864). (PI. 15, Figs 7, 8;
specimen diameter 120 ^m)

Cambridge Barbadoes Johnson 1862, BM 2770; Cambridge Barbadoes Johnson, BM 2806;
Cambridge Barbadoes Johnson 1863, BM 2868; Cambridge Barbadoes Johnson 1864, BM 3069;
Barbadoes Johnson 1863, BM 3090; Barbadoes 1865, BM 3116; Cambridge Barbadoes Johnson
1864, BM 3123; Barbadoes 1865, BM 3474.

BM 3116 and BM 3069 (PI. 15, fig. 5; specimen diameter 185 /am) are the only slides that have
been annotated by Greville as A. decorus. The remaining slides were all annotated at a later date
by Rattray. However, Rattray has annotated BM 3116 and BM 3069 as A. decorus var. stoschii.
It seems probable that one of these two slides was used by Rattray to describe his variety and his
synonymy has to be doubted on nomenclatural grounds alone. Additionally, some of the
specimens on other slides are quite dissimilar to those illustrated by Greville. To establish the
distinctness of A. decorus sensu Rattray, type specimens of A. stoschii need to be examined.

In the detailed study of Audacodiscus by Burke & Woodward, BM 3116 is indicated by Mr R.
Ross as being a good representative specimen (1967: 161). They discuss this species along with
the related A. inflatus, A. mammosus, and A. cinctus.

6. A. extans Grev. in Trans, microsc. Soc. Lond. 12: 87, fig. 1 (1864). (PI. 15, Figs 1, 2; specimen
diameter 212-5

GREVILLE DIATOM TYPES 15

Barbadoes 1865, BM 3109, holotype.

7. A. gigas Grev. in Trans, microsc. Soc. Lond. 13: 26, fig. 23 (1865). (PI. 15, Fig. 4; specimen
diameter 210 /am)

Cambridge Barbadoes Johnson 1863, BM 2903, holotype.

8. A. grevilleanus G. Norman in Grev. in Trans, microsc. Soc. Lond. 12: 10, fig. 1 (1864).
Moron, Seville, BM 2692, holotype.

I was unable to find the specimen at the MF location given on the slide.

9. A. inflatus Grev. in Trans, microsc. Soc. Lond. 11: 69, fig. 12 (1863). (PI. 16, Figs 1, 2, 4;
specimen diameter 82-5 /am)

Barbadoes deposit Johnson 1863, BM 2841, holotype.

10. A. jonesianus Grev. in Trans, microsc. Soc. Lond. 10: 24, fig. 5 (1862). (PI. 15, Fig. 3;
specimen diameter 267-5 /urn)

Guano Dr Macrae, BM 2388, holotype.

11. A. kilkellyanus Grev. in Trans, microsc. Soc. Lond. 11: 70, fig. 14 (1863). (PI. 16, Fig. 5;
specimen diameter 82-5 /am)

Barbadoes deposit Johnson 1863, BM 2804, holotype.

12. A. macraeanus Grev. in Trans, microsc. Soc. Lond. 10: 23, fig. 4 (1862). (PI. 16, Fig. 3;
specimen diameter 135 /am)

Ceylon Dr Macrae, BM 2387, lectotype, BM 2450; New Zealand, BM 1683.
There are two slides annotated for this species and, as there is no drawing, either could be the
holotype. They are both selected mounts. BM 2387 has been chosen as lectotype. Additionally,
Greville refers to a New Zealand slide. This specimen is present on BM 1683.

13. A. mammosus Grev. in Trans, microsc. Soc. Lond. 11: 70, fig. 13 (1863). (PI. 16, Figs 7, 8;
specimen diameter 96-5 /am)

Barbadoes Johnson 1863, BM 2813, holotype.

14. A. orientalis Grev. in Trans, microsc. Soc. Lond. 12: 12, fig. 6 (1864). (PI. 17, Figs 1-3;
specimen diameter 142-5 /am)

Ceylon 1861, BM 2425; Ceylon Dr Macrae 1861, BM 2448, BM 2419, lectotype; Ceylon Dr
Macrae, BM 2466, BM 3394.

There is a drawing in the Greville manuscript collection but no indication of which slide was used
for the illustration. Any of the above slides could have been used and must be considered as
syntypes. BM 2419 (MF 32/25) has been selected as lectotype.

15. A. pallidus Grev. in Trans, microsc. Soc. Lond. 11: 72, fig. 17 (1863).
Barbadoes deposit, BM 2827, holotype.

This is a badly damaged slide and hence it is not possible to present micrographs. Burke (pers.
comm. P. A. Sims) has examined the holotype specimen and established several important facts.
The specimen is a frustule, rather than a single valve. Greville had not realized this and judged
the specimen to have ten processes when there are only five on each valve. Rattray apparently
also noticed this fact (Rattray, 1888ft: 370).

16. A? paradoxus Grev. in Trans, microsc. Soc. Lond. 11: 72, fig. 18 (1863). (PI. 16, Fig. 6;
specimen diameter 85 /urn)

Barbadoes deposit Johnson 1863, BM 2783, holotype.

In the tenth paper in the 'New and rare' series, Greville (1863d) published a corrigendum
appearing opposite plate IX. He states that this species should have been referred to the genus
Omphalopelta. The slide is labelled ' Omphalopelta' '.

17. A. pellucidus Grev. in Trans, microsc. Soc. Lond. 12: 12, fig. 5 (1864). (PI. 17, Figs 4, 5;
specimen diameter 110 /am)

[Cambridge Estate Barbadoes Johnson], BM 2893, holotype.

16 D. M. WILLIAMS

18. A. radiatus Grev. in Trans, microsc. Soc. Lond. 12: 11, fig. 4 (1864). (Not illustrated)
Cambridge Barbadoes Johnson 1864, BM 3125; Cambridge Barbados Johnson 1863, BM 2898,
syntypes.

19. A. sparsus Grev. in Trans, microsc. Soc. Lond. 14: 123, fig. 6 (1866). (PI. 17, Fig. 6;
specimen diameter 72-5 /mi)

Barbadoes 1865, BM 3479, holotype.

20. A. spectabilis Grev. in Trans, microsc. Soc. Lond. 11: 71, fig. 16 (1863). (PI. 18, Figs 1, 2;
specimen diameter 132-5 /mi)

Barbadoes Johnson 1863, BM 2837, holotype.

21. A. umbonatus Grev. in Trans, microsc. Soc. Lond. 12: 9, fig. 2 (1864). (PI. 17, Fig. 7;
specimen diameter 92-5 /mi)

Cambridge Estate Barbadoes Johnson 1863, BM 3051, lectotype; Cambridge Estate Barbadoes
Johnson 1862, BM 2774.

There is a drawing in the Greville manuscript collection but no indication of which slide he used
for the illustration. Either of the above could have been used. BM 3051 has been chosen as
lectotype. Although it is a fragment, there is less debris obscuring the valve than that of BM
2774. Additionally, some ink has spilt on to the label of BM 2774, obscuring part of the notes.

XII. Al/L/SCI/SEhrenb.

1. A. ambiguus Grev. in Trans, microsc. Soc. Lond. 11: 74, fig. 23 (1863). (PI. 18, Figs 3, 4;
specimen diameter 57-5 /mi)

Barbadoes Johnson, BM 2842; Barbadoes Johnson 1862, BM 2815, syntypes.
The coverslip of BM 2815 has probably moved at some time as the specimen can no longer be
located at the MF number given on the label by Greville. It is also a badly damaged slide.
Greville notes in the protologue: 'I have examined two specimens and a fragment of a third all of
which are precisely alike.' The second specimen, BM 2842, has been used for the micrographs.

2. A. australiensis Grev. in Trans, hot. Soc. Edinb. 8: 233, fig. 3 (1866).
'Sharks Bay, Norman 810' (MS notes, but slide not found).

The drawing in the Greville manuscript collection is annotated thus: 'Shark's Bay - Norman's
slides (810 19/17).' This slide does not appear to be in the Greville collection or elsewhere in the
BM. The whereabouts of these Norman slides needs further investigation. There is a series of
manuscript notes confirming the Norman slide as the holotype. Rattray (1888a: 869) had
probably seen this slide as he cites the locality data and notes, 'Formerly in the Collection of Mr.
George Norman.'

3. A. barbadensis Grev. in Trans, microsc. Soc. Lond. 13: 5, fig. 1 (1865). (PI. 18, Fig. 5;
specimen length 50 /mi)

Cambridge Barbadoes Johnson 1864, BM 3206, holotype.

4. A. cellulatus Grev. inJIR. microsc. Soc. 1888: 904 (1888), in synonymy. (PI. 18, Figs 7, 8;
specimen diameter 65 /mi)

Barbadoes Johnson, BM 3128, BM 67371, [Ralfs colln.]

Greville's name was never published. Rattray included this species in the synonymy of
Pseudauliscus ralfsianus (Grev.) Rattray. Only two slides in the BM collection reveal specimens
of this taxon: BM 3128 and BM 67371 from Ralfs' collection. The micrograph is from BM 3128.

5. A. elaboratus Ralfs ex Grev. in Trans, microsc. Soc. Lond. 11: 51, fig. 19 (1863). (PI. 18, Fig.
6; specimen diameter 100 /mi)

Cambridge Barbadoes Mr Johnson, BM 66995, holotype?; Cambridge Barbadoes Johnson

1862, BM 2786; Cambridge Barbadoes Johnson 1862, BM 2802; Barbadoes Johnson 1862, BM

2818; Cambridge Barbadoes Johnson 1863, BM 3055; Cambridge Barbadoes Johnson, BM

3273.

There is no drawing in the Greville manuscript collection. There are many slides in the Greville

GREVILLE DIATOM TYPES 17

collection which could be considered syntypes. However, in the protologue Greville states:
'kindly communicated by Mr. Ralfs.' It is thus possible that the holotype is a Ralfs' slide. There
are two specimens on a Ralfs slide, BM 66995, but only one, MF 28/16, resembles Greville's
illustration and hence has been photographed.

6. A. elegans Grev. in Trans, microsc. Soc. Lond. 11: 45, fig. 8 (1863). (PI. 19, Figs 1, 4;
specimen diameter 72-5 /xm)

Patos Guano Johnson, BM 3007 (MF 24/12), lectotype.

There is a drawing of this species in the Greville manuscript collection but two MF numbers are
indicated. Of the two, the specimen located at MF 24/12 is less obscured by debris and has been
photographed here as the lectotype.

7. A. hardmanianus Grev. in Trans, microsc. Soc. Lond. 14: 6, fig. 17 (1866). (PI. 19, Fig. 3;
specimen diameter 137-5 /xm)

Monterey, BM 10108, L.H. no. 139, holotype.

8. A. johnsonianus Grev. in Trans, microsc. Soc. Lond. 11: 51, fig. 20 (1863). (PI. 19, Fig. 5;
specimen diameter 80 /Am)

Cambridge Estate Barbadoes 1863, BM 3065, holotype.

9. A. macraeanus Grev. in Trans, microsc. Soc. Lond. 11: 51, fig. 18 (1863). (PI. 19, Fig. 7; PI.
20, Fig. 1; specimen diameter 97-5 /tin)

Ceylon Dr Macrae 1861, BM 2409, holotype.

10. A. mirabilis Grev. in Trans, microsc. Soc. Lond. 11: 47, fig. 11 (1863).
Monterey Stone, Br. 11 (slide not found).

Greville made the following remarks in the protologue: 'It was first bought under my notice by
Mr. Norman, but his specimen had unfortunately sustained some superficial injury. That
subsequently supplied by Mr. Browne leaves nothing to be desired. My own specimen is a
fragment.'

Additionally, the drawing indicates that the slide supplied by Mr Browne was used for the
illustration, with the following annotation: 'Br. 11, 24/32.' This slide has not been located in the
Greville collection or elsewhere in the BM. It's whereabouts requires further investigation.
There are no slides in the Greville collection purported to be A. mirabilis. The only suitable
lectotype, if one need be required, is in the Hardman collection, BM 10130, which has a label
that states: 'Named by Dr Greville.'

11. A. moronensis Grev. in Trans, microsc. Soc. Lond. 12: 83, fig. 6 (1864). (PI. 19, Fig. 6;
specimen diameter 63 /Am)

Moron deposit Seville Johnson 1863, BM 2913, BM 2916.

Although there is a drawing in the Greville manuscript collection annotated as 'Norman 451',
I have not been able to trace this slide. Two other slides are present and both are regarded
as syntypes. This species has been examined in detail by Sullivan (1987), who concludes
A. moronensis is a synonym of A. pruinosus J. W. Bail.

12. A. nebulosus Grev. in Trans, microsc. Soc. Lond. 11: 74, fig. 21 (1863). (PI. 19, Fig. 8; PI. 20,
Fig. 2; specimen diameter 85 /am)

Cambridge Estate Barbadoes Johnson 1863, BM 2898, lectotype, BM 3061; Cambridge Estate
Barbadoes Johnson 1862, BM 2821.
The correct name for this species is Pseudauliscus grevillei (see appendix).

13. A. normanianus Grev. in Trans, microsc. Soc. Lond. 12: 82, fig. 11 (1864). (PI. 20, Fig. 3;
specimen diameter 112-5 /am)

Moron deposit Johnson 1863, BM 2915, BM 2917, lectotype, BM 2919, BM 2920.

There is a drawing of this species in the Greville manuscript collection but it is not annotated.

BM 2917 has been chosen as the lectotype.

14. A. notatus Grev. in Trans, microsc. Soc. Lond. 13: 5, fig. 2 (1865). (PI. 20, Fig. 4; specimen
diameter 37 -5 /Am)

18 D. M. WILLIAMS

[Cambridge Estate Barbadoes Johnson], BM 3138, holotype.

This slide has suffered some damage, hence the poor quality of the micrograph.

15. A. ornatus Grev. in Trans, microsc. Soc. Lond. 12: 88, fig. 2 (1864). (PI. 20, Figs 6, 7;
specimen diameter 55 /am)

Cambridge Barbadoes 1864, BM 3038, holotype.

16. A. parvulus Grev. in Trans, microsc. Soc. Lond. 11: 74, fig. 22 (1863). (PI. 20, Fig. 5;
specimen diameter 34-5 /am)

Cambridge Barbadoes Johnson 1863, BM 2862, holotype.

17. A. racemosus Ralfs ex Grev. in Trans, microsc. Soc. Lond. 11: 46, fig. 9 (1863). (PI. 21, Figs
1, 4, 5; specimen diameter 62 /am)

Barbadoes Cambridge Johnson [Ralfs colln.], BM 66853, holotype?; Barbadoes, Cambridge
Johnson 1862, BM 2786; Barbadoes, Johnson, 1862, BM 2790; Barbadoes, Cambridge,
Johnson, 1863, BM 2902.

Although there is a drawing of this species in the Greville manuscript collection, there are no
slide details appended. The protologue states that the slide was 'Communicated by J. Ralfs,
Esq.' and hence Ralfs' slide is probably the holotype.

18. A. ralfsianus Grev. in Trans, microsc. Soc. Lond. 11: 52, fig. 21 (1863). (PI. 21, Fig. 6;
specimen diameter 102-5 /am)

Bridgewater Barbadoes Johnson 1863, BM 2737, holotypes.

19. A. recticulatus Grev. in Trans, microsc. Soc. Lond. 11: 46, fig. 10 (1863). (PI. 21, Figs 2, 3;
specimen diameter 89-5 /am)

From Melobesia on Haliotis tuberculata, coast of Peru, BM 19772 (MF 20/21), holotype.
Greville noted two localities in the protologue: 'Cape of Good Hope, G. M. Browne. From
Melobesia on Haliotis tuberculata, Peru, F. C. S. Roper, Esq.' The drawing in the Greville
manuscript collection is marked with the Roper locality and hence must be considered the
holotype. The slide is in the Roper collection in the BM.

XIII. BERKELEYA Grev.

This genus has recently been studied in detail by Cox (1975a, b) and the generic definition
emended. The type species, B. fragilis, has also been examined in both the light and electron
microscope (see Cox, 1975a, fig. 6; 1975b, fig. 10, also p. 214 for an amended description).

1. B. fragilis Grev. in Scott. Cryptog. Fl. 5: pi. 298 (1827). (PI. 21, Fig. 7; specimen length 65

/am)

On Zostera marina, Appin, Capt. Carmichael, BM 77941, BM 81243, BM 81244, BM 81245,

Holotype, ex K.

This material forms part of the permanent loan from Kew. The slides were recently made from

the herbarium sheets. The micrograph is from BM 77941.

XIV. BIDDULPHIAGray

1. B. barbadensis Grev. in Trans, hot. Soc. Edinb. 8: 437, fig. 15 (1866). (PI. 22, Figs 7-9; valve
length 25 /am, frustule width 65 /am)

Barbadoes 1866, BM 3480, holotype.

Greville states: The frustule now figured is the only one which has come under my notice.'
Although this is not identified to species on the slide label, it is quite clearly that of Greville's
figure and hence is the holotype. Additionally, a set of manuscript notes clearly identify this
specimen as the holotype.

2. B. chinensis Grev. in Trans, microsc. Soc. Lond. 14: 81, fig. 16 (1866). (PI. 25, Fig. 3;
specimen length 206-5 /am)

Hong Kong Harbour, BM 2998.

There is a drawing in the Greville manuscript collection but no slide number is written on it. BM

GREVILLE DIATOM TYPES 19

2998 is the only slide catalogued for this species, of which there are two specimens indicated by
MF numbers. The micrograph is the best specimen of the two. The mountant has drastically
receded and renders photography difficult.

3. B. corpulenta Grev. in Trans, microsc. Soc. Lond. 13: 51, fig. 16 (1865). (PI. 22, Fig. 6;
specimen length 155 /xm)

Cambridge Barbadoes /o/wson 1863, BM 3078, holotype.

The specimen is situated on a damaged part of the slide and hence difficult to photograph. Much

of the structure is reduced in clarity as a consequence.

4. B? decorata Grev. in Trans, microsc. Soc. Lond. 13: 99, fig. 7 (1865). (PI. 21, Fig. 8; specimen
length 66 /urn)

Cambridge Barbadoes Johnson 1864, BM 3432, holotype; Cambridge Barbadoes 1865, BM
3447.

There are no drawings of this species in the Greville manuscript collection. There are two
specimens in the slide collection and in the protologue Greville states: 'I have only seen two
specimens of this diatom', hence both must be considered as types. However, the specimen on
BM 3447 is situated obliquely, whilst that of BM 3432 is identical to the published figure and
hence is the holotype. According to Mr R. Ross, careful examination of the holotype shows that
it is a broken specimen of a tripolar diatom.

5. B. elegantula Grev. in Trans, microsc. Soc. Lond. 13: 50, figs 12-14 (1865). (PI. 22, Figs 4, 5;
specimen length 51-5 /xm)

Cambridge Estate Barbadoes Johnson 1863, BM 3073; Barbadoes Johnson 1864, BM 3412;
Cambridge Barbadoes Johnson 1865, BM 3434, syntypes.

There are four drawings in the Greville manuscript collection but only three were published.
The drawing for figures 12 and 13 are on one card and annotated as [BM] 3073, 14/29. This
specimen corresponds to figure 12. The second card has two drawings, one of a valve view and
one of a girdle view, but only the valve view was used in the publication. The annotation on this
card is [BM] 3412 which corresponds to the unpublished girdle view. The originals of figures 13
and 14 have not been located.

6. B. fimbriata Grev. in Trans, microsc. Soc. Lond. 13: 6, fig. 4 (1865). (PI. 22, Figs 1, 2; valve
length 92-5 /xm)

Barbadoes Johnson 1864, BM 3197, lectotype; Cambridge Barbadoes Johnson 1863, BM 3186.
There are two drawings in the Greville manuscript collection but no indication is given as to
which slides they are taken from. Greville also notes that he has seen 'half a dozen specimens.'
The specimen on BM 3197 has several girdle bands and is closer to the published figure than the
other specimens.

7. B. gigantea Grev. in Trans, microsc. Soc. Lond. 12: 13, fig. 9 (1864). (PI. 23, Figs 1, 2;
specimen length 205 /u,m)

Cambridge Barbadoes Johnson 1862, BM 2770, holotype.

8. B. inflata Grev. in Trans, microsc. Soc. Lond. 13: 50, fig. 15 (1865). (PI. 23, Figs 3, 4; valve
length 88 /xm)

Barbadoes Johnson 1864, BM 3428, holotype.

9. B. johnsoniana Grev. in Trans, microsc. Soc. Lond. 14: 6, figs 14, 15 (1866). (PI. 22, Fig. 3, PI.
23, Figs 5,6; valve length 170 /mi)

Moron deposit Seville Johnson 1865, BM 3396, syntypes.

There are two drawings included in the original publication and two drawings in the Greville
manuscript collection. Both of these specimens are present on the same slide. Mr R. Ross has
inspected these specimens and confirmed that they are those that Greville illustrated and hence
are syntypes.

10. B. longicruris Grev. in Q. Jl microsc. Sci. 7: 163, fig. 10 (1859). (PI. 23, Figs 7-9; specimen
length 32-5 /mi)

20 D. M. WILLIAMS

Californ. guano/. T. Norman 58, BM 1773, lectotype; Californ. guano/. T. Norman, BM 1770;
Californ. guano/. T. Norman 8.58, BM 1820.

11. B? mammosa Grev. in Trans, microsc. Soc. Lond. 14: 7, fig. 16 (1866). (PI. 24, Figs 1, 2;
specimen length 96-5 /am)

Barbadoes 1865, BM 3455, holotype.

12. B. minutissima Grev. in Trans, hot. Soc. Edinb. 8: 437, fig. 14 (1866). (PI. 24, Fig. 3; valve
length 12-5 /am, frustule length 31 /am)

Zanzibar Prof. H. L. Smith 1865, BM 3562, lectotype, BM 3563, BM 3564.

13. B. nitida Grev. in Trans, microsc. Soc. Lond. 13: 52, fig. 11 (1865). (PI. 24, Figs 4, 6; valve
length 48-5 /am, frustule length 82-5 /am)

Barbadoes, BM 3254, holotype?

14. B? podagrosa Grev. in Trans, microsc. Soc. Lond. 14: 82, fig. 17 (1866). (PI. 24, Figs 5, 8;
valve length 65 /am, frustule length 98-5 /am)

Barbadoes Johnson 1863, BM 3054; Cambridge Barbadoes Johnson 1863, BM 3072.

In the Greville manuscript collection there is a drawing annotated with two numbers. These

specimens should be considered syntypes. However, there are some manuscript notes which

cite only BM 3054. It is likely that this was used for the description rather than BM 3072.

The type slides have previously been studied by Wise (1952: 407) and further study was

undertaken recently in connection with some new Biddulphioid genera by Ross & Sims (1985:

317).

15. B. punctata Grev. in Trans, microsc. Soc. Lond. 12: 83, fig. 10 (1864). (PI. 24, Fig. 7;
specimen length 120 /am)

Cambridge Barbadoes Johnson 1863, BM 2894, holotype.

16. B. roperiana Grev. in Q. Jl microsc. Sci. 7: 163, figs 11-13 (1859). (BM 1763, PI. 25, Figs 1,
2; specimen length 67-5 /am; BM 1772, PI. 24, Figs 9, 10; specimen length 38-5 /am)
Monterey Sea weed [G.] Norman 12.58, BM 1632; Patos guano 8.58 /. T. Norman, BM 1755;
Patos guano 8.58, BM 1758, BM 1818; Californ. guano, BM 1762, BM 1765, BM 1782;
[Californ. guano], BM 1763, BM 1776; Californ. guano/. T. Norman 9.58, BM 1768; Californ.
guano/. T. Norman 58, BM 1772, BM 1797, syntypes.

Two localities were given in the protologue: Calif ornian guano and Monterey. The Monterey
seaweed sample was not used for the illustrations. Greville included three figures of this species
with the description but no drawings exist in the manuscript collection. All the above slides are
syntypes. An investigation of the slides helps reveal which specimens were used for the
illustrations. BM 1763 was most probably the specimen in figure 11; BM 1797 (not illustrated)
was the specimen in figure 12; and BM 1772 was the specimen in figure 13.

Later Greville stated: 'I am now convinced that my B. roperiana is nothing more than one of
the endless varieties of B. aurita.' (Greville, 1866c: 81.)

17. B. sinuata Grev. in Trans, microsc. Soc. Lond. 13: 49, fig. 10 (1865).
Barbadoes Johnson 1862, BM2815, holotype.

Although all the relevant information is available on the drawing in Greville 's manuscripts,
there is a note in the slide collection which states: 'Slide and coverslip smashed to atoms. A. G.
[Anthony Gepp] February 1915.' The specimen cannot be located and photographed.

18. B. spinosa Grev. in Trans, microsc. Soc. Lond. 13: 6, fig. 3 (1865). (PI. 25, Fig. 4; specimen
length 73-5 /am)

Barbadoes 1865, BM 3114, holotype?; Cambridge Barbadoes Johnson 1864, BM 3125.
There are two drawings in the Greville manuscript collection, annotated with the above two
slide numbers. These are the only slides Greville referred to and hence must be considered
syntypes. However, BM 3125 is not orientated at the same angle as that of the illustration. It is
likely that BM 3114 was used for the published figure. This specimen has been used for the
micrograph.

GREVILLE DIATOM TYPES 21

19. B. tenuicornis Grev. in Trans, microsc. Soc. Lond. 13: 51, fig. 17 (1865). (PI. 25, Figs 5, 6;

specimen length 68-5 /am)

Barbadoes Cambridge Estate Johnson 1862, BM 2777, holotype.

XV. BRIGHTWELLIA Ralfs
This genus has recently been reviewed by Andrews (1986).

1. B. elaborata Grev. in Trans, microsc. Soc. Lond. 9: 73, fig. 1 (1861). (PI. 25, Figs 7, 10;
specimen diameter 73-5 /am, central part of valve only illustrated)

Barbadoes Johnson 1863, BM 2099, holotype.

2. B. johnsonii ['Joknsonf] Ralfs ex Grev. in Trans, microsc. Soc. Lond. 14: 4, fig. 11 (1866).
(PI. 25, Figs 8, 9; specimen diameter 91-5 /am)

Springfield Barbadoes Hardman 1864, BM 3341, isotype.

The protologue for this species mentions both Cambridge and Springfield estates for the locality
('most abundant in the latter.') There is a drawing in the Greville manuscript collection which is
marked 'Barbadoes, Springfield H(21)', suggesting that Greville used a Hardman slide for his
illustration. The slide in the Greville collection is probably isotype material and has been used
for the micrographs. Additionally, two slides in the Hardman collection are considered isotypes,
one of which may be the holotype: BM 10159 and BM 10160.

XVI. CAMPYLODISCUSEhrenb. ex Kiitz.

1. C. ambiguus Grev. in Trans, microsc. Soc. Lond. 8: 31, fig. 5 (1860). (PI. 26, Figs 1, 2;
specimen length 65-5 /am)

Jamaica, BM 1708, lectotype, BM 1711, BM 1835.

Two localities are given in the protologue: 'In washings of small algae from Jamaica' and 'On
Spondyli (fragment) in a slide communicated by Mr. George Norman' . The lectotype is from the
former locality.

2. C. biangulatus Grev. in Trans, microsc. Soc. Lond. 10: 20, fig. 2 (1862). (PI. 26, Figs 3, 4;
specimen length 105-5 /am)

Ceylon Dr Macrae, BM 2385, lectotype, BM 2421, BM 2467.

3. C. browneanus Grev. in Trans, microsc. Soc. Lond. 10: 89, fig. 2 (1862).
Manilla Shell scrapings G. M. Browne, BM 2382, BM 4250.

These two slides are the only ones from the type locality collected by G. M. Browne. However,
on neither slide is there a specimen of C. browneanus and the type slide remains to be
discovered.

4. C. crebrecostatus Grev. in Trans, microsc. Soc. Lond. 11: 14, fig. 6 (1863).
'Port Jackson, New South Wales; Dr. Roberts.'

There is no drawing or slide indicated in the Greville collections. In the protologue Greville
states: 'In a mounted slide presented to me by Dr. Roberts.' This slide has not been located.

5. C. diplostictus G. Norman ex Grev. in Trans, microsc. Soc. Lond. 8: 31, fig. 6 (1860). (PI. 26,
Figs 5,6; specimen length 182-5 /am)

Shark Bay Norman, BM 1946, lectotype; Ascidians Sharks Bay Australia (Dr MacDonald)
Norman (213), 59, BM 1948.

6. C. ecclesianus Grev. in Q. Jl microsc. Sci. 5: 10, fig. 5 (1857). (PI. 26, Figs 7, 8; specimen
length 86-5 /am)

Trinidad Mrs Eccles, BM 999, holotype?

No drawing exists in Greville's manuscripts for this species. This specimen is the only one which

corresponds to the published figure.

7. C. fenestratus Grev. in Q. Jl microsc. Sci. 5: 9, fig. 4 (1857). (PI. 26, Fig. 9; specimen length
120 /am)

22 D. M. WILLIAMS

Trinidad Mrs Eccles 9.55, BM 995, holotype?

No drawing exists in Greville's manuscripts for this species. This specimen is the only one which

corresponds to the published figure.

8. C. humilis Grev. in Trans, hot. Soc. Edinb. 8: 233, fig. 1 (1866). (PI. 26, Fig. 10; specimen
length 24-5 /xm)

[Woodlark Island Dr Roberts], BM 2729, holotype.

9. C. imperialis Grev. in Trans, microsc. Soc. Lond. 8: 30, fig. 3 (1860). (PI. 26, Fig. 11;
specimen length 135 /zm)

Nassau W. Ind., BM 1744, lectotype; Shell scrapings Nassau W. Ind., BM 1732.
Greville states in the protologue that: 'In one slide only does a perfect example of this species
now figured occur.' A check of the relevant slides revealed the specimen Greville referred to on
slide BM 1744. This has been designated as lectotype.

10. C. kittonianus Grev. in Trans, microsc. Soc. Lond. 8: 32, fig. 7 (1860). (PI. 27, Figs 1, 2;
specimen length 142-5 /xm)

On Tridacna W. Indies Kitton 59, BM 1947 (MF 26/27), lectotype.

Greville gave two localities in the protologue. Slides of the second, 'On Spondyli; George

Norman, Esq', have only fragments present.

11. C. normanianus Grev. in Trans, microsc. Soc. Lond. 8: 29, fig. 1 (1860).
'In cleanings of Spondyli; George Norman, Esq.'

There is no drawing or slide indicated in the Greville collections.

12. C. notatus Grev. in Trans, microsc. Soc. Lond. 8: 31, fig. 4 (1860).
'In cleanings of Spondyli; George Norman, Esq.'

There is no drawing or slide indicated in the Greville collections.

13. C. ornatus Grev. in Trans, microsc. Soc. Lond. 11: 13, fig. 3 (1863). (Not illustrated)
On Tridacna W. Indies Kitton, BM 2567.

There is a drawing in the Greville manuscript collection of this species, but no information is
given as to which slide was used for the description.

14. C. robertsianus Grev. in Trans, microsc. Soc. Lond. 11: 14, fig. 5 (1863). (PI. 27, Figs 3, 4;
specimen length 97-5 /am)

Harvey Bay Queensland Dr Roberts 1862, BM 2674, holotype.

The drawing in the manuscript collection is annotated with 'Harvey Bay 10, 30/32'. The only

slide which has a specimen at those MF co-ordinates is the above and hence is considered the

holotype.

15. C. stellatus Grev. in Q. Jl microsc. Sci. 7: 157, fig. 3 (1859). (PI. 27, Figs 5, 6; specimen
length 102-5 /urn)

Californian guano/. T. Norman 8.58, BM 1779, holotype.

16. C. undulatus Grev. in Q. Jl microsc. Sci. II, 3: 229, fig. 4 (1863). (PI. 26, Fig. 12; specimen
length 88-5 /Am)

[Bermuda in mud brought up on a fluke of an anchor G. Norman], BM 2695, holotype?, BM

2694.

In the Greville manuscript collection there is a drawing with two numbers. The numbers refer to

two slides which have blank labels but using the MF co-ordinates two specimens were located.

BM 2695 is probably the holotype (MF 15/26). Additional material is present on slides BM 4847

and BM 4859.

17. C. wallichianus Grev. in Trans, microsc. Soc. Lond. 11: 13, fig. 4 (1863). (PI. 27, Figs 7, 8;
specimen length 102-5 /am)

Dredged off St. Helena Dr Wallich, BM 3951, lectotype; Harvey Bay Dr Roberts 1862, BM

2676; Off St. Helena Wallich, BM 3950; St. Helena, BM 3966.

There is no drawing of this species in the Greville manuscript collection. The slides indicated

GREVILLE DIATOM TYPES 23

above are those that correspond to the type localities given and marked by Greville for this
species.

XVII. CESTODISCUSGrev.

1. C. johnsonianus Grev. in Trans, microsc. Soc. Lond, 13: 48, fig. 8 (1865). (PI. 27, Figs 10, 11;
specimen diameter 51-5 /urn)

Moron deposit Seville Johnson 1865, BM 3396, holotype?

No drawing exists in Greville's manuscripts for this species. Greville states in the protologue:
'extremely rare, Mr. Johnson and myself have only found one specimen each.' The above slide
has one specimen, the other specimen is located on BM 3238.

2. C. (?) ovalis Grev. in Trans, microsc. Soc. Lond. 13: 49, fig. 9 (1862). (PI. 27, Fig. 9; specimen
length 101 -5 /Am)

Moron deposit Seville Hardman 1865, BM 3238; Moron deposit Seville Rev. T. G. Stokes 1866,

BM3235.

There is a drawing of this species in the Greville manuscript collection but there is no number

indicating a slide. Although BM 3235 is attributed to Rev. T. G. Stokes as collector, no specimen

is indicated on the label and the date is later than that of the published species. A micrograph has

been included from BM 3238, but this is unlikely to be type material.

3. C. pulchellus Grev. in Trans, microsc. Soc. Lond. 14: 123, fig. 5 (1866).
Nankaurie deposit, Nicobar Islands, G. Norman.

There is a drawing in the Greville manuscript collection that is annotated 'Nankaurie, 26/39' but
no slide number is given. The holotype specimen has yet to be located.

4. C. stokesianus Grev. in Trans, microsc. Soc. Lond. 14: 123, fig. 4 (1866).
Moron deposit Seville Rev. T. G. Stokes, BM 3235, holotype?

No drawing exists in Greville's manuscript collection for this species, nor has any specimen been
located in the Greville slide collection.

XVIII. CLADOGRAMMAEhrenb.

1. C. conicum Grev. in Trans, microsc. Soc. Lond. 13: 97, figs 1-2 (1865). (Not illustrated)

Cambridge Estate Barbadoes Johnson 1865, BM 3443, BM 3448.

There is one drawing in the Greville manuscript collection annotated with two numbers. There is

also an extra drawing which was apparently not published. BM 3443 (MF 27/15) corresponds to

Greville's figure 1, whilst BM 3448 (MF 30/35) corresponds to Greville's figure 2. These must

both be regarded as syntypes. The specimens are situated at difficult angles for any useful

photography.

XIX. CLAVULARIAGrev.

1. C. barbadensis Grev. in Trans, microsc. Soc. Lond. 13: 24, figs 1-3 (1865). (BM 3080, PI. 28,
Figs 8, 10; specimen length 161-5 /xm; BM 3273, PI. 28, Figs 7, 9; specimen length 137-5 /xm)
Barbadoes Cambridge Estate 1863, BM 3273; Barbadoes Johnson 1863, BM 3080; BM 2863,
3049, 3076, 3080, 3086, 3093, 3095, 3103, 3135, 3144, 3177, 3183, 3206, 3214, 3243, 3246, 3259,
3267, 3273, 3275, 3280, 3282, 3411, 3419, 3421, 3429, 3430, 3432, 3438, 3442, 3448, 3461, 3463,
3474, 3507, 3511, syntypes.

There is a drawing of this species in the Greville manuscript collection but no slide numbers to
identify those used for the figures.

XX. COCCONEISEhrenb.

1. C. armata Grev. in Trans, microsc. Soc. Lond. 14: 126, fig. 13 (1866). (PI. 28, Fig. 6;
specimen length 47-5 /u,m)
Barbadoes 1866, BM 3480, holotype.

24 D. M. WILLIAMS

2. C. arraniensis Grev. in Q. Jl microsc. Sci. 7: 80, fig. 2 (1859). (PI. 29, Figs 1-3; specimen
length 36-5 /xm)

Arran 57, BM 1560, holotype?

There is no drawing in the Greville manuscript collection but as this is the only slide catalogued

for this species, it is assumed to be the holotype.

3. C. barbadensis Grev. in Trans, microsc. Soc. Lond. 12: 14, fig. 10 (1864). (PI. 29, Figs 5, 6;
specimen length 86 /am)

Barbadoes/o/zrawi, BM 2751, holotype.

4. C. coelata Arn. ex Grev. in Q. Jl microsc. Sci. II, 2: 234, figs 5-6 (1862). (PI. 29, Fig. 4;
specimen length 32-5 /xm)

Eupleuria ocellata on Ballia, New Zealand Walker- Arnott 56, (S. 631), BM 1492, syntypes.
There is a drawing in the Greville manuscript collection but it is not annotated with a slide
number. However, this is the only slide in the collection of Arnott's material and is probably the
holotype slide. There are numerous specimens on the slide, which should all be considered
syntypes.

5. C. crebrestriata Grev. in Q. Jl microsc. Sci. 5: 9, fig. 2 (1857). (PI. 29, Figs 7, 8; specimen
length 59 /mi).

Trinidad Eccles 9.56, BM 997, lectotype; Trinidad Mrs Eccles 9.55, BM 995.

BM 997 has been chosen as lectotype because the alternative specimen, on BM 995, is partially

obscured by debris.

6. C. grantiana Grev. in Trans, microsc. Soc. Lond. 9: 72, fig. 18 (1861). (PI. 29, Fig. 9;
specimen length 27-5 /xm)

Macduff on sea shells Mr Grant, BM 2244, syntypes.

There is a drawing in the Greville manuscript collection of this species but no indication as to
which slide is the holotype. The only slide that corresponds to the locality details is the one given
above. As it contains many specimens they must all be considered syntypes. The slide is in poor
condition and only allows effective use at low magnification.

7. C. granulifera Grev. in Trans, microsc. Soc. Lond. 9: 73, fig. 19 (1861). (PI. 29, Fig. 10;
specimen length 36-5 /u,m)

On Clam shells Carrickfergus Mr Grant, BM 2243, syntypes.

There is a drawing in the Greville manuscript collection of this species but no indication is given
as to which slide is the holotype. BM 2243 is the only slide from the locality given in the
protologue. As it contains many specimens they are all considered syntypes. The slide is in a
poor condition and only allows effective use at low magnification.

8. C. inconspicua Grev. in Q. Jl microsc. Sci. 5: 9, fig. 3 (1857).
Trinidad Mrs W. Eccles.

There is no indication either in the manuscript notes or the slide collection which slide Greville
used for the published illustration.

9. C. naviculoides Grev. in Trans, microsc. Soc. Lond. 13: 34, fig. 24 (1865). (PI. 29, Figs 11, 12;
specimen length 60-5 /xm)

Barbadoes, Johnson 1863, BM 2845, holotype?

This is the only slide in the Greville collection which has a specimen of this taxon and therefore is
probably the holotype. The slide is unfortunately damaged and it is only possible to examine it at
low magnification.

10. C. punctatissima Grev. in Q. Jl microsc. Sci. 5: 8, fig. 1 (1857). (PI. 29, Figs 13, 14; specimen
length 45 ^tm)

Trinidad 56, BM 994, holotype?

This is the only slide in the collection that is annotated for this species, although many other

slides from the same locality are present. The slide is rather thick which prevents the use of high

magnification.

GREVILLE DIATOM TYPES 25

11. C. regalis Grev. in Q. Jlmicrosc. Sci. 7: 156, fig. 1 (1859). (PI. 30, Figs 6, 7; specimen length

65-5 /am)

Calif orn. guano/. T. Norman. BM 1777, holotype?

This is the only slide in the collection from the type locality and therefore is probably the

holotype. Several other slides are marked by Greville for specimens of C. regalis but none are

from the type locality.

XXI. COSCINODISCUS Ehrenb.

Coscinodiscus , a large marine plankton genus, is currently being revised (Hasle & Sims, 1986).
Many of Greville's species will need to be re-allocated to other genera. Several species have
already been transferred to Azpeitia M. Perag. (Fryxell, Sims & Watkins, 1986).

1. C. angulatus Grev. in Trans, microsc. Soc. Lond. 12: 9, fig. 11 (1864). (PI. 30, Fig. 3;
specimen diameter 70 /am)

Cambridge Barbadoes Johnson 1862, BM 2825, holotype.

2. C. armatus Grev. in Trans, microsc. Soc. Lond. 9: 42, fig. 5 (1861). (PI. 30, Figs 4, 5;
specimen diameter 52-5 /am)

Barbadoes deposit, BM 2080, lectotype, BM 2082, BM 2168.

The plate legend states that the Barbados specimens illustrating this paper were described from
a series of slides supplied by Mr J. T. Norman. I have not been able to trace these slides and
hence a lectotype has been selected from Greville's slide collection.

3. C. barbadensis Grev. in Trans, microsc. Soc. Lond. 9: 43, fig. 9 (1861).
Barbadoes deposit, BM 2072, BM 2076, BM 2077, syntypes.

The plate legend states that the Barbados specimens illustrating this paper were described from
a series of slides supplied by Mr J. T. Norman. These slides have not been traced. A lectotype
has not been designated.

4. C. biradiatus Grev. in Trans, microsc. Soc. Lond. 9: 42, fig. 7 (1861). (PI. 31, Fig. 1; specimen
diameter 80 /am)

Barbadoes deposit, BM 2064, BM 2067, BM 2082 lectotype, BM 2111, BM 2181.
The plate legend states that the Barbados specimens illustrating this paper were described from
a series of slides supplied by Mr J. T. Norman. I have not been able to trace these slides and
hence a lectotype has been selected from Greville's collection.

This species has recently been studied in detail and transferred to the genus Azpeitia (Fryxell,
Sims & Watkins, 1986).

5. C. elegans (Grev. in Trans, microsc. Soc. Lond. 14: 3, fig. 6 (1866). (PI. 31, Figs 2, 4;
specimen diameter 96-5 /am)

Monterey, BM 10221, L. H. 281, holotype.

The type specimens of Coscinodiscus elegans and Cosmiodiscus elegans share the same slide,
BM 10221, causing some confusion. The two species are quite distinct but unfortunately the MF
number given for Cosmiodiscus elegans, if applied in normal use, i.e. left hand side of the slide
down, does not reveal any specimen. The slide needs to be reversed for the MF number to locate
the specimen. The type of Cosmiodiscus elegans is a solitary specimen, whereas that of
Coscinodiscus elegans is placed within a cluster of specimens. The drawing for Coscinodiscus
elegans is marked 'H. 49 Monterey' and the slide is marked in the right hand corner with a
diamond marker inscription as L.H.49.

6. C. elegantulus Grev. in Trans, microsc. Soc. Lond. 9: 42, fig. 8 (1861). (PI. 31, Fig. 3;
specimen diameter 42-5 /am)

Barbadoes deposit, BM 2064, BM 2069, BM 2075 lectotype, BM 2076, BM 2078, BM 2080, BM
2081, BM 2085, BM 2096, BM 2098.

The plate legend states that the Barbados specimens from this paper were described from a
series of slides supplied by Mr J. T. Norman. I have not been able to trace these slides and hence
a lectotype has been selected from Greville's slide collection.

26 D. M. WILLIAMS

This species has been placed in the genus Azpeitia by Sims in Fryxell, Sims & Watkins (1986).

7. C. excavatus Grev. ex Ralfs in Pritch., Hist. Inf. : 829 (1861). (PI. 32, Figs 3, 6, 7: specimen
diameter 245 /xm)

Piscataway Earth Feb. 59, BM 67530 (Ralfs colln), holotype?; Piscataway U.S. deposit Bailey
Dallas, BM 1307.

Although this species bears a Greville name it was not published until Ralfs' account in the Hist.
Inf. It is more than likely the holotype slide was that of Ralfs' own collection. Only one slide in
Greville's collection is from Piscataway, the type locality. Other specimens are from Barbados,
Cambridge estate (BM 3137, BM 3143) and South Naparima, Trinidad (BM 3820).

8. C. griseus Grev. in Q. Jl microsc. Sci. II, 3: 230, fig. 7 (1863). (PI. 31, Fig. 6; specimen
diameter 89 -5 /Am)

Cambridge Barbadoes Johnson 1862, BM 2772, holotype.

The drawing in the manuscript collection has both a slide number and a MF location. Greville

had not included the species name on the slide label.

9. C. lewisianus Grev. in Trans, microsc. Soc. Lond. 14: 78, figs 8-10 (1866). (BM 1304, PI. 31,
Fig. 7; specimen diameter 53-5 /urn; BM 1305, PI. 31, Fig. 9; specimen diameter 89-5 /xm)
Rappahannock U.S. 56, BM 1304, lectotype, BM 1305; Rappahannock Virginia, Dallas, BM
1306.

There is no drawing of this species in the Greville manuscript collection. However, there are
some notes Greville made whilst writing the paper which indicate that the above slides were
examined. One of these has been selected as a lectotype.

10. C. macraeanus Grev. in Trans, microsc. Soc. Lond. 13: 46, fig. 4 (1865). (PI. 31, Figs 5, 8;
specimen diameter 128-5 /xm)

Guano Dr Macrae, BM 2444, holotype.

11. C. moronensis Grev. ex Rattray in Proc. R. Soc. Edinb. 16: 458, pi. II, fig. 16 (1890). (PI. 30,
Figs 1,2; specimen diameter 73-5 /urn)

Moron deposit Seville Johnson 1864, BM 3348, holotype.

Rattray used a Greville manuscript name for his description of C. moronensis. Greville,
however, wanted to place the species in the genus Cestodiscus . In his published figure, Rattray
erroneously refers to Johnson as the authority on the plate legend.

12. C. mossianus Grev. in Trans, microsc. Soc. Lond. 13: 25, fig. 22 (1865). (PI. 32, Fig. 1;
specimen diameter 130-5 /urn)

Cambridge Barbadoes Johnson 1864, BM 3248, holotype?

Although there is no drawing in the Greville manuscript collection, this is the only slide that has

a specimen of this taxon and is probably the holotype.

13. C. obliquus Grev. ex Rattray in Proc. R. Soc. Edinb. 16: 575, pi. 1, fig. 14 (1890). (PI. 32,
Figs 2, 4, 5; specimen diameter 171-5 ptm)

Monterey Stone, BM 1971.

Although Rattray gives two localities for this species, it is clear that he based his description and
the name on the manuscript notes of Greville. Greville had originally referred this species of
Cosmiodiscus .

14. C. oblongus Grev. in Trans, microsc. Soc. Lond. 14: 4, figs 9, 10 (1866). (PI. 28, Figs 1-4;
specimen diameter for Fig. 1, 82-5 /am, Fig. 2, 80-5 /xm, Fig. 3, 83-5 /urn, Fig. 4, 80-5 /urn)
Barbadoes Springfield Estate BM 10192, L.H. 263, BM 10208 L.H. 262, isotypes.

The drawing in the Greville collection is marked 'Barbad. Springfield H(3)'. The two figures are
from the above two slides. All the examples of this species in Greville's collection are annotated
by Rattray and not identified to species by Greville. The Hardman slides have many selected
individuals. The micrographs have been taken from BM 10192.

This species was transferred to Craspedodiscus by Hanna (1931) as Craspedodiscus oblongus
(Grev.) Hanna, which is a later homonym of Craspedodiscus oblongus (Grev.) Grun. in A.

GREVILLE DIATOM TYPES 27

Schmidt (1881), based on Porodiscus oblongus Grev. Combos (1982) recognized that Hanna's
name was illegitimate and published the new name Craspedodiscus ellipticus Gombos for this
species, erroneously calling it a comb, nov., not a nom. nov.

15. C. patellaeformis Grev. in Trans, microsc. Soc. Lond. 9: 80, fig. 4 (1861). (PI. 31, Fig. 10;
specimen diameter 77-5 /am)

Barbadoes Earth, BM 2178, holotype.

16. C. pulchellus Grev. in Trans, microsc. Soc. Lond. 14: 3, fig. 7 (1866). (PI. 33, Fig. 1;
specimen diameter 140 /am)

Barbadoes Cambridge Estate Johnson 1864, BM 3424, holotype.

17. C. robustus Grev. in Trans, microsc. Soc. Lond. 14: 3, fig. 8 (1866). (PI. 28, Fig. 5; specimen
diameter 195 /am)

Monterey, BM 10831 L.H. no. 1073, holotype.

The protologue states that the specimens came from the 'cabinet of L. Hardman', hence the

above slide is the holotype.

18. C. scintillans Grev. in Q. Jl microsc. Sci. II, 3: 230, fig. 6 (1863).
'Barbadoes deposit, from Cambridge estate.' Slide not found.

The slides used for the description of this taxon came from Mr G. M. Browne. They have not
been located.

19. C. splendidus Grev. in Trans, microsc. Soc. Lond. 13: 44, fig. 3 (1865). (PI. 33, Figs 2, 5, 6;
specimen diameter 91-5 /am)

Barbadoes Cambridge Estate Johnson 1863, BM 3110, syntypes.

The slide above is that which Greville indicates on the drawing in the manuscript collection.
There are two MF locations, 23/23, and 27/14. These have been identified as syntypes by Mr R.
Ross. Specimen 27/14 has been used for the micrographs.

20. C. symmetricus Grev. in Trans, microsc. Soc. Lond. 9: 68, fig. 2 (1861). (PI. 34, Figs 6-8;
specimen diameter 55 /am)

Barbadoes deposit, BM 2076, holotype.

21. C. tuberculatus Grev. in Trans, microsc. Soc. Lond. 9: 42, fig. 6 (1861). (PI. 34, Figs 1, 2;
specimen diameter 55 /am)

Barbadoes deposit, BM 2063, BM 2069, BM 2082, syntypes.

These slides are syntypes. The micrographs have been taken from BM 2063 (MF 33/13). This
species has recently been studied by Sims (in Fryxell, Sims & Watkins, 1986) and transferred to
the genus Azpeitia.

XXII. COSMIODlSCUSGrev.

Simonsen (1974) has suggested that the Greville species will probably be found to possess a
pseudonodulus and thus belong in the Hemidiscaceae. The status of the genera in the
Hemidiscaceae is uncertain and requires revision.

1. C. barbadensis Grev. in Trans, microsc. Soc. Lond. 14: 80, fig. 12 (1866). (PI. 34, Figs 3-5;
specimen diameter 86-5 /am)

Cambridge Barbadoes Johnson 1864, BM 3027, holotype.

2. C. elegans Grev. in Trans, microsc. Soc. Lond. 14: 79, fig. 13 (1866). (PI. 33, Figs 3, 4;
specimen diameter 90-5 /am)

Monterey deposit, BM 10221, L.H. no. 281.
See notes under Coscinodiscus elegans.

Rattay wanted to transfer this species to Coscinodiscus, but as C. elegans was preoccupied by
Greville's earlier name, Rattray proposed a new name: Coscinodiscus perikompsis (Rattray
1890: 576, pi. Ill, fig. 12).

3. C. normanianus Grev. in Trans, microsc. Soc. Lond. 14: 80, fig. 11 (1866).

28 D. M. WILLIAMS

'Barbadoes deposit; cabinet of George Norman, Esq.' Slide not found.

The slide used by Greville for the published illustration of this species came from George

Norman. Norman's slides have not been traced.

Rattray (1890) had wished to transfer this species into Coscinodiscus but considered the
combination Coscinodiscus normanianus undesirable as the name was too similar to that of
Coscinodiscus normanii Greg, in Grev. and hence he proposed a new name, Coscinodiscus
apages Rattray. As pointed out by VanLandingham (1968: 860), Rattray's reasons are in-
adequate . Furthermore, Coscinodiscus normanii has been transferred to Actinocyclus (Hustedt,
1957: 218). Rattray does not list a Greville specimen in his protologue.

Greville erroneously referred his specimen to Coscinodiscus in the caption to the figure.

XXIII. CRASPEDODISCUS Ehrenb .

1. C. umbonatus Grev. in Trans, microsc. Soc. Lond. 14: 79, fig. 15 (1866). (PI. 33, Fig. 7;
specimen diameter 85 /xm)
Barbadoes 1865, BM 3469, holotype.

XXIV. CRASPEDOPORUSGrev.

This genus has recently been revised by Hendey & Sims (1987), including a re-evaluation of the
Greville species.

1. C. johnsonianus Grev. in Trans, microsc. Soc. Lond. 11: 69, fig. 10 (1863). (Illustrated in
Hendey & Sims, 1987: 32, fig. 35)

Cambridge Estate Barbadoes Johnson, BM 2769, holotype.

2. C. ralfsianus Grev. in Trans, microsc. Soc. Lond. 11: 68, fig. 9 (1863). (PI. 34, Fig. 9;
specimen diameter 92-5 /u,m)

Cambridge Estate Barbadoes Johnson 1863, BM 2742, lectotype.

A drawing is present in the Greville manuscript collection but no details given of the slide used
for the illustration. A series of manuscript notes state that Greville consulted a Ralfs' slide. This
slide has not been traced. Hendey & Sims (1987) have designated BM 2742 as lectotype.

XXV. CRESWELLIA Arn. ex Grev.

1. C. annulata Grev. in Ann. Mag. nat. Hist. Ill, 16: 4, fig. 8 (1865). (PI. 34, Figs 10, 11;
specimen diameter 26-5 /x,m)

Hong Kong Harbour 14, BM 4511, holotype?

In the protologue Greville indicates that he had seen several specimens: 'I have repeatedly seen
two in connexion and have at the present moment a chain of three under my eye.' The above
specimen is therefore considered the holotype. The mountant of the slide is of poor quality and
hence the micrographs are not of considerable use.

2. C. barbadensis Grev. in Trans, microsc. Soc. Lond. 13: 3, fig. 11 (1865). (PI. 35, Figs 1, 2;
specimen length 45 /am)

Cambridge Estate Barbadoes Johnson, BM 3190, BM 3248; Barbadoes 1864, BM 3163;
Barbadoes 1864, BM 3457; Barbadoes, 1865, BM 3245, lectotype.

There is a drawing in the Greville manuscript collection but with no annotations. Greville states
that he had seen many specimens, but none of the above slides match the published illustration.
It is possible Greville may have composed his illustration from all the valves he examined.

3. C. cylindracea Grev. in Trans, microsc. Soc. Lond. 13: 3, fig. 10 (1865).
Hong Kong Harbour/. L. Palmer 1863, BM 3229.

The specimen Greville chose to illustrate is one with four linked frustules. The above slide is the
only one labelled for this species. However, no specimen has been located at the given MF
number.

GREVILLE DIATOM TYPES 29

4. C? ferox Grev. in Q. Jl microsc. Sci. 7: 166, figs 15, 16 (1859). (PI. 35, Figs 3, 4; specimen
diameter 48-5 /mi)

Californ. Guano/. T. Norman 8.58, BM 1772, BM 1779; Californ. Guano BM 1777, BM 1782.
As Greville gave two figures of this species, the originals of both are syntypes. Figure 15 is of two
frustules, and figure 16 is a single valve. Greville states: 'I have only twice had an opportunity of
examining them in connexion'. BM 1782 is marked 'Creswellia ferox (united)', but is of two
valves. The other slides are of single girdle views.

5. C. ininuta Grev. in Trans, microsc. Soc. Lond. 13: 4, fig. 13 (1865). (PI. 35, Fig. 5; specimen
length 15-5 /xm)

Barbadoes Johnson 1864, BM 3185, holotype.

This slide is of poor quality and hence only low magnification micrographs are possible.

6. C. palmeriana Grev. in Trans, microsc. Soc. Lond. 13: 2, fig. 9 (1865). (PI. 35, Figs 7, 8;
specimen diameter 57-5 /Ltm)

[Hong Kong Habour /. L. Palmer] 12, BM 3232; Shark's Bay west coast of Australia in
Ascidians Dr MacDonald, BM 1948.

There is a drawing in the Greville manuscript collection but it is not annotated with a slide
number. The best example of this species was, according to Greville, on a L. Hardman slide.
This slide has not been traced. Two localities were given in the protologue. The first of the two
slides given above has no locality details on the slide label. However, from information on
neighbouring slides and comparison with the species composition, it is probably from Hong
Kong. Greville indicated on the label that this species was present. The second slide corresponds
to the Shark's Bay locality but the label is not annotated for this species. The Shark's Bay
material is of valve and girdle views.

7. C. rudis Grev. in Trans, microsc. Soc. Lond. 14: 78, fig. 7 (1866). (PI. 35, Fig. 6; specimen
diameter 97- 5 /^m)

Monterey Earth L. Hardman 1865, BM 3352, BM 3232, syntypes.

The drawing in the Greville manuscript collection is annotated with two numbers: '[BM] 3352'
and 'Monterey 54H'. The Hardman slide is of a girdle view. Due to the state of BM 3352 and the
condition of the mountant, the specimen is difficult to photograph and little detail is revealed.
Micrographs have been taken from BM 3232.

8. C. sphaerica Grev. in Trans, microsc. Soc. Lond. 13: 4, fig. 12 (1865).
Cambridge Barbadoes, Johnson 1862, BM 2821, holotype?

Greville illustrates a specimen with several inter-connected frustules: 'Although very rare, I
have seen, at least, a score of specimens, but only a single example of frustules in connexion.'
The above slide purportedly corresponds to the drawing and must be considered as the holotype.
However, no specimen was found.

9. C. superba Grev. in Trans, microsc. Soc. Lond. 9: 68, figs 3-5 (1861). (BM 2022, PI. 36, Figs
1, 2; specimen diameter 108-5 /am; BM 2015 (MF 28/14), PI. 36, Fig. 3; specimen diameter 60-5
/xm)

Barbadoes deposit, BM 2015, BM 2022, syntypes.

In the Greville manuscript collections there are three drawings, each of which corresponds to a
published figure. The drawings are annotated with two slide numbers. BM 2015 has two MF
locations, 16/32 and 28/14, which correspond to figures 3 (not illustrated) and 4 respectively. BM
2022 has a single specimen marked at 31/28 and is also annotated by Greville as 'large', which
corresponds to figure 5. These must be considered syntypes.

10. C. turgida Grev. in Q. Jl microsc. Sci. 7: 165, fig. 14 (1859). (PI. 36, Figs 4-6; specimen
diameter 38-5 /xm)

Californ. Guano, BM 1777, lectotype.

There are many slides in the Greville collection that could be the holotype for this species but
without manuscript evidence it is impossible to decide. BM 1777 was selected as lectotype as the
specimen is a whole frustule.

30 D. M. WILLIAMS

11. C. turns Grev. & Arn. in Greg, in Trans. R. Soc. Edinb. 21: 538, pi. 14, fig. 109 (1857).

(PI. 36, Fig. 9; specimen diameter 55-5 /am, specimen length 210 /am)

Teignmouth Arnott, from Ascidians, BM 47; Teignmouth Arnott, 1857, BM 1057; Ascidia,

Cumbrae Arnott (740), BM 2268, syntypes.

Greville gave two localities in the protologue for this species. However, the illustration is of four

interlocked frustules which corresponds to the specimen on BM 47. Greville states: 'Island of

Cumbrae, where it was dredged with the nests of Lima hians by Mr Kennedy, and a single

f rustule detected by Dr Walker- Arnott . ' BM 2268 is the slide of Walker- Arnott's single f rustule .

XXVI. CYMATOPLEVRA^f . Sm.

1. C. angulata Grev. in Trans, microsc. Soc. Lond. 10: 89, fig. 1 (1862). (PI. 37, Fig. 9; specimen

length 82-5 /AHI)

Patos guano 8.58/. T. Norman, BM 1756, lectotype, BM 1757; Californ. guano, BM 1765, BM

1777.

XXVII. CYMBELLA Agardh

1. C. lindsayana Grev. in Trans, hot. Soc. Edinb. 8: 234, figs 5-8 (1866). (PI. 36, Figs 7, 8;

specimen length 40 /zm)

Otago New Zeald. Dr. Lauder Lindsay, 1861 (15), BM 3549, syntypes.

The drawing in the Greville manuscript collection is not annotated. This is the only slide in the

collection that corresponds to the locality data given and hence must be considered type

material. There are two MF location annotations on the slide label but no indication as to which

species they refer. Both co-ordinates reveal specimens of C. lindsayana. MF 25/24 has been used

for the micrograph. Lindsay (1865) published a detailed account of the collecting sites for these

New Zealand diatoms. This species was included in his checklist as a nom. nud. , indicating that

Greville was to describe them later.

XXVIII. DIATOMAEory

1. D. brachygonium Grev. in Hook., Brit. Fl. 5: 406 (1833).

[Appin], Marine, Captain Carmichael.

This species has not been traced in the Greville herbarium.

XXIX. DIATOMELLA Grev.

A recent revision of this genus has revealed much about the submicroscopic details of its valve
and girdle morphology. D. balfouriana, the type of the genus, was not studied (LeCohu, 1983).

1. D. balfouriana Grev. in Ann. Mag. nat. Hist. II, 15: 259, figs 10-13 (1855); in Trans, hot. Soc.

Edinb. 5: 51, pi. IV, figs 10-13 (1855). (PI. 37, Figs 1, 2; specimen length 22-5 pm)

Glen Callater, 8.54 (127), BM 561; Braemer Balfour 9.54, BM 563; Glen Tilt, 8.55, BM 726,

syntypes.

Several localities are noted in the protologue. All the above slides are syntypes. This species was

originally named, but not published, by W. Smith as Grammatophora balfouriana. The

micrographs are of a ringed specimen on BM 563.

XXX. DJCLAD/AEhrenb.

Ross & Sims (1985) studied Greville's species and showed that they belong to the genus
Didadiopsis De Toni. They also showed that D. robusta Grev. is a synonym of D. barbadensis
Grev.

1. D? barbadensis Grev. in Trans, microsc. Soc. Lond. 13: 56, fig. 28 (1865). (PI. 37, Fig. 3;

specimen length 130 /xm)

Cambridge Estate Barbadoes Johnson 1864, BM 3416, holotype.

GREVILLE DIATOM TYPES 31

Three figures from the type slide are given in Ross & Sims (1985). Plate 31, fig. a is the holotype
specimen, while figs b and c are additional specimens from the holotype slide.

2. D? robusta Grev. in Trans, microsc. Soc. Lond. 13: 98, fig. 11 (1865). (PI. 37, Fig. 6;

specimen length 122-5 /u,m)

Barbadoes Johnson 1864, BM 3179, holotype.

XXXI. DICTYOPYXISEhrenb.

1. D. brevis Grev. in Trans, microsc. Soc. Lond. 10: 22, fig. 2 (1862). (PI. 37, Figs 4, 5; specimen

length 40-5 /am)

Ceylon Dr Macrae, BM 2390, lectotype; Andaman Islds Dr Macrae 1861, BM 2434.

The published illustration is of two frustules plus a detached valve. Neither of the above slides

has such an arrangement. Greville states that he had only seen two examples of this taxon. These

may be the above slides. BM 2434 is of several united frustules but does not match those given in

the drawing.

XXXII. ECH/JVELLAAgardh

1. E. circularis Grev. in Trans. Wernerian Soc. 4: 213, pi. 8, fig. 2 (1822).
Rivulet near Dumbryden Quarries, 1820, BM 81187.

The slide is made from material that is on permanent loan from Kew. It is stored dried on to mica
and mounted in envelopes on to herbarium sheets. The slide was made from a small portion of
this material. This species has been studied in detail by Williams (1985: 158-172).

2. E. fasciculata [|3] truncata Grev. in Scott. Crypt. Fl. 1: pi. 16, fig. 4 (1822).

'Aquae dulcis. On various Confervae and on all aquatic plants about Edinburgh, March 1821,
leg Greville\ BM 81237, BM 81238, BM 81250.

These slides have been made from material that is on permanent loan from Kew. The valves
have fragmented and no whole specimens were found.

XXXIII. ENTOGONIAGrev.

This genus has been recently studied in detail by Holmes & Brigger (1977, 1979) who, along with
Bergon (1892a, b), had access to the Greville types. Many of the species Greville described
under Triceratium are now placed in this genus.

1. E. amabilis Grev. in Q. Jl microsc. Sci. II, 3: 236, fig. 21 (1863). (PI. 37, Fig. 7; distance across
one side 96-5 /xm)

Cambridge Barbadoes Johnson 1863, BM 2874, holotype.

There is a drawing in the Greville manuscript collection annotated with the above slide number.
The specimen is found at MF 25/21. However, the glass of the slide is thick and does not allow
high magnification to be used.

2. E. conspicua Grev. in Q. Jl microsc. Sci. II, 3: 236, fig. 23 (1863).
'Barbadoes deposit, from Cambridge estate; G. Norman, Esq.'

There is a drawing in the Greville collection which is annotated with 'no. 15, Norman'. There is
no slide in Greville's collection that corresponds to this particular number. Additionally, there
are no slides in the Greville collection indexed for this species. The only slide in the BM of this
species is in the Hardman collection. No lectotype has been designated until it is certain that the
Norman slides cannot be found.

3. E. elegans Grev. in Trans, microsc. Soc. Lond. 13: 33, fig. 19 (1865).
'Barbadoes deposit, Cambridge estate; C. Johnson, Esq.'

There are no slides in the Greville collection indexed for this taxon. Bergon (18926) reported the
absence of the type. It is yet to be found.

32 D. M. WILLIAMS

4. E. punctulata Grev. in Q. Jlmicrosc. Sci. II, 3: 237, fig. 24 (1863).
'Barbadoes deposit, from Cambridge estate; G. Norman, Esq.'

There is a drawing in the Greville collection which is annotated with 'Norman'. There is no slide
of this species in the Greville collection. The only slide in the BM of this species is in the
Hardman collection. No lectotype has been designated until it is certain that the Norman slides
cannot be found.

5. E. reticulata Grev. in Trans, microsc. Soc. Lond. 12: 85, fig. 5 (1864). (PI. 37, Fig. 8; distance
across one side 82-5 /am)

Cambridge Barbadoes Johnson July 1863, BM 2901, holotype.

The specimen is at an acute angle on the slide, presenting difficulty for accurate photography.

6. E. venulosa Grev. in Q. Jlmicrosc. Sci. II, 3: 236, fig. 22 (1863).
'Barbadoes deposit, from Cambridge estate; G. Norman, Esq.'

There is a drawing in the Greville collection which is annotated with 'Norman, no. 14'. There is
no slide in the Greville collection that corresponds to this particular number. Additionally, there
are no slides in the Greville collection indexed for this species. The slide prepared by Meakin,
BM 36179 from Joes River, Barbados, has been chosen as neotype. See also Barker & Meakin
(1944: 19) and Holmes & Brigger (1979: 194).

XXXIV. EUODIA J. W. Bail, ex Ralfs

1. E. harbadensis Grev. in Trans, microsc. Soc. Lond. 9: 69, figs 6, 7 (1861). (PI. 37, Fig. 10;

specimen length 30-5 /am)

Barbadoes deposit, BM 2075, BM 2081, syntypes.

The drawing in the Greville manuscript collection corresponding to figure 6 is BM 2075 (MF

18/14, illustrated). The specimen in figure 7 corresponds to BM 2081 (MF 15/39). These

specimens are therefore syntypes.

XXXV. EUPODISCUS Ehrenb .

1. E. barbadensis Grev. in Trans, microsc. Soc. Lond. 12: 88, fig. 4 (1864). (PI. 38, Fig. 1;
specimen diameter 55-5 jam)

Barbadoes Johnson 1865, BM 3104, holotype.

2. E. hardmanianus Grev. in Trans, microsc. Soc. Lond. 14: 80, fig. 14 (1866). (PI. 38, Figs 2, 3;
specimen diameter 127-5 /am)

From shell cleanings So. America L. Hardman 1866, BM 3492, holotype?

A drawing is available in the Greville manuscript collection but no slide details are given. There

is only one slide in Greville's collection.

3. E. jonesianus Grev. in Trans, microsc. Soc. Lond. 10: 22, fig. 3 (1862). (PI. 38, Figs 4-6;
specimen diameter 153-5 /am)

[Guano Unknown locality] Dr Macrae, BM 2389, holotype?

A drawing is available in the Greville manuscript collection but no slide details are given. There

is only one slide in Greville's collection.

4. E. minutus Grev. in Trans, microsc. Soc. Lond. 14: 5, fig. 13 (1866).
'Barbadoes deposit, Springfield estate; cabinet of L. Hardman, Esq.'

The drawing in the Greville manuscript collection is annotated 'Barb. Springfield H(4)' referring
to a Hardman slide. A footnote in Rattray (1888a: 911) states: 'I am informed by Mr. Hardman
that the original of the species is lost, the slide on which it was mounted having been broken in
the Post Office.' There is no appropriate neotype in the collection and such a designation should
properly await a revision of the species.

5. E. obscurus Grev. in Trans, microsc. Soc. Lond. 10: 90, fig. 4 (1862).
'Cape of Good Hope; Dr. Macrae.'

This slide has not been found, nor is it in Greville's collection.

GREVILLE DIATOM TYPES 33

6. E. oculatus Grev. in Trans, microsc. Soc. Lond. 10: 90, fig. 3 (1862).

Monterey Stone Johnson 1862, BM 2623 (MF 24/15); Monterey Stone Johnson (Arnott), BM
3015.

In the protologue Greville cites George Norman and Fred R. Kitton as sources of the material.
However, material from these sources is not present in the Greville collection. The two slides,
indicated above, are the only ones which purport to have examples of this taxon. The specimen
on BM 2623 (MF 24/15) does not appear to be close to that which Greville illustrated and is
marked as ' Eupodiscus ocellatus Grev.' and may have originally been intended as a different
species. A lectotype has not been designated until this specimen can be studied further.

7. E. parvulus Grev. ex Rattray in Trans. R. microsc. Soc. 1888: 912, pi. 14, fig. 8 (1888). (PI.
38, Figs 7, 8; specimen diameter 52-5 /mi)

Barbadoes, Johnson, BM 2751, lectotype; Barbadoes deposit, Johnson, BM 2769; Barbadoes,
Johnson, 1862, BM 2756, BM 2771, BM 2794, BM 2826; Cambridge, Barbadoes, Johnson,

1862, BM 2753, BM 2774, BM 2784, BM 2816, BM 2825; Cambridge, Barbadoes, Johnson,

1863, BM 2873, BM 3057, BM 3102; Cambridge, Barbadoes, Johnson, 1864, BM 3420;
Barbadoes, Johnson, 1865, BM 3449; Barbadoes, 1865, BM 2451.

8. E. punctulatus Grev. in Trans, microsc. Soc. Lond. 11: 73, fig. 19 (1863). (PI. 38, Fig. 10;
specimen diameter 77-5 /mi)

Cambridge Barbadoes Johnson 1863, BM 2752, holotype.

9. E. scaber Grev. in Trans, microsc. Soc. Lond. 12: 81, fig. 1 (1864). (PI. 38, Fig. 11; specimen
diameter 180-5 /mi)

Barbadoes Johnson 1863, BM2943, holotype?

There is a drawing in the Greville manuscript collection but no details of the slide used are given.

The slide above is the only one in the collection of this species from the type locality.

10. E. simplex Grev. in Trans, microsc. Soc. Lond. 11: 73, fig. 20 (1863). (PI. 38, Fig. 9;
specimen diameter 90-5 /mi)

Cambridge Barbadoes Johnson 1862, BM 2770, holotype.

11. E. trioculatus Grev. in Trans, microsc. Soc. Lond. 12: 88, fig. 3 (1864). (PI. 39, Fig. 1;
specimen diameter 77 /mi)

Cambridge Estate Barbadoes Johnson 1863, BM 3050, BM 3047, BM 3048.
A drawing in the Greville collection is annotated with the above three slide numbers but no
reference to MF locations. Greville also states that: 'three specimens, of which I have before
me'. These three must be considered syntypes. However, BM 3050 is a poor slide and BM 3047 is
catalogued as missing.

XXXVI. EXILARIA Grev.

1. E. fulgens Grev. in Scott. Crypt. Fl. 5: pi. 291, figs 1, 2 (1827).

'On various small algae in the sea, in spring and summer. Appin, Captain CARMICHAEL.'

No material is present in BM.

XXXVII. FENESTRELLA Grev.

A revision, with an expanded generic description, is given in Swatman (1948). Swatman

examined Greville's type slide, but did not include micrographs of Greville's material in his

paper.

1. F. barbadensis Grev. in Trans, microsc. Soc. Lond. 11: 68, fig. 8 (1863). (PI. 39, Fig. 2;

specimen diameter 108 /mi)

Barbadoes Johnson 1863, BM 2813, holotype.

34 D. M. WILLIAMS

XXXVIII. GEPHYRIAArn.

1. G. constricta Grev. in Trans, microsc. Soc. Land. 14: 77, fig. 2 (1866). (PI. 39, Fig. 3;
specimen length 163 /un)

Monterey deposit L. H. no. 336, BM 10266, holotype.

2. G. gigantea Grev. in Trans, microsc. Soc. Lond. 14: 122, figs 7, 8 (1866).
Monterey L. Hardman, BM 3360, holotype?

The two published illustrations are present in Greville's manuscript collection but there are no
annotations indicating the slides from which they were taken. In a set of manuscript notes there
are further comments: 'F. V. lower valve no. Upper Valve no. 76.' This may refer to a Hardman
slide. BM 3360 is a Hardman slide but it is in Greville's own collection. The valve does not
resemble either of the published figures.

XXXIX. GLYPHODESMISGrev.

1. G. eximia Grev. in Q. Jl microsc. Sci. II, 2: 235, figs 7-10 (1862). (PI. 39, Figs 10, 11;

specimen length 35 /xm)

Jamaica in washings of small algae, BM 1708; Nassau Norman, BM 1741, syntypes.

In the protologue Greville states: 'my drawings of this species from the West Indian specimens.'

These two slides are considered syntype material. This species has been studied in detail by

Sullivan (in press).

XL. GLYPHODISCUSGrev.

1. G. stellatus Grev. in Trans, microsc. Soc. Lond. 10: 91, fig. 5 (1862). (PI. 39, Figs 4, 5;

specimen diameter 28-5 /zm)

Monterey Stone, Kitton, BM 3542, BM 3543, BM 3586; Monterey Stone, BM 2575, BM 3158,

BM 3588, BM 3589, BM 3590, syntypes.

There are many examples of this taxon in the Greville collection. Greville states: The most

perfect example which I have seen, and which I have represented, is in Mr. George Norman's

cabinet.' It is not clear whether Greville retained this slide and where, if he did not, Norman's

slide is. It is not in the BM. The above slides are syntypes. The micrograph is taken from BM

3542.

This species has been examined recently in LM by Bone (1969) and in the SEM by Stidolph
(1985).

XLI. GONIOTHECIUMEhrenb.

1. G. prolongatum Grev. in Trans, microsc. Soc. Lond. 13: 56, fig. 29 (1865). (PI. 39, Figs 6, 7)
Barbadoes Johnson 1864, BM 3415, syntypes.

The slide above indicates two specimens, either of which could be the holotype. It is difficult to
tell them apart and hence both are regarded as syntypes (PI. 39, Fig. 6, MF 16/20, specimen
length 107-5 /am; PI. 39, Fig. 7, MF 17/26, specimen length 95 //-m).

XLII. GRAMMATOPHORA Ehrenb.

1. G. moronensis Grev. in Q. Jl microsc. Sci. II, 3: 229, fig. 5 (1863). (PI. 39, Fig. 9; specimen
length 52-5 /zm)

Moron deposit Spain 1862, BM 2905, holotype.

2. G. pusilla Grev. in Trans, hot. Soc. Edinb. 7: 574, fig. 15 (1863); in Edinb. New phil. J. 18:
181, fig. 15 (1863). (PI. 39, Fig. 8; specimen length 37-5 /am)

Curteis Straits Queensland Dr Roberts 1863, BM 2598, lectotype, BM 2624.
There is a drawing of this species in the Greville manuscript collection but no slide numbers are
indicated. The illustration Greville published is of two frustules. Neither of the above slides
correspond exactly to the drawings, which may have been part of the entire range of valves
Greville studied.

GREVILLE DIATOM TYPES 35

XLIH. HEIBERGIA Grev.

1. H. barbadensis Grev. in Trans, microsc. Soc. Lond. 13: 100, figs 8-9 (1865). (BM 3102, PI.
40, Figs 1, 2; specimen length 135-5 /um; BM 2773, PI. 40, Fig. 3; specimen length 97-5 /urn)
Barbadoes 1865, BM 3102; Cambridge Barbadoes /o/wsort 1862, BM 2773, syntypes.
The drawings in the Greville manuscript collection are annotated with two slide numbers. BM
3102 (MF 27/13) is probably figure 9 and BM 2773 (MF 32/15) is probably figure 8.

XLIV. HELIOPELTA Ehrenb.

1. H. nitida Grev. in Trans, microsc. Soc. Lond. 14: 5, fig. 18 (1866). (PI. 40, Figs 4, 5; specimen

diameter 79 yum)

Los Angeles deposit California L. Hardman 1865, BM 3239, holotype?

There is a drawing in Greville's manuscript collection but no details are given of the slide

number. This is the only slide in Greville's collection that has specimens of H. nitida. There is a

group of four arranged amongst other selected species.

XLV. HEMIA UL US Heiberg, nom. cons. prop.

Greville included several species in Hemiaulus temporarily, recognizing that they should be
placed elsewhere, possibly in new genera. In particular, H. tenuicornis, H. lyriformis, H.
longicornis, H. alatus, H. hastatus, H. ornithocephalus, and H. capitatus were dealt with in this
way. Recently Ross & Sims (1985) removed the latter two species, placing them in a new genus,
Briggera R. Ross & P. A. Sims. The remaining Greville species were initially transferred into
Reidelia Jouse & Sheshukova-Poretskaya by Schrader & Fenner (1976). However, there is
continued confusion over the exact identity of Reidelia, and these species remain, at present, in
Hemiaulus.

1. H. alatus Grev. in Trans, microsc. Soc. Lond. 13: 31, fig. 14 (1865). (PI. 40, Fig. 6; specimen
length 38-5 /u-m, height from base to tips, 48-5 /u,m)

Barbadoes Johnson 1865, BM 3439.

There is only one slide in the collection referable to this species, but no MF location is provided.

However, a specimen was discovered at MF 17/21.

2. H. angustus Grev. in Trans, microsc. Soc. Lond. 13: 30, fig. 12 (1865). (PI. 40, Figs 7-9;
specimen length 30-5 /urn, height from base to tips, 117-5 /am)

Barbadoes Johnson 1864, BM 3426 lectotype, BM 3427; Barbadoes, 1865, BM 3105.

3. H?? capitatus Grev. in Trans, microsc. Soc. Lond. 13: 54, fig. 24 (1865). (PI. 40, Figs 11, 12;
specimen length 37-5 /u-m, height from base to tips, 35 /u-m)

Barbadoes Johnson 1864, BM 3426, holotype.

For a full discussion of this species and its relationships see Ross (1972) and Ross & Sims (1985:

303-305).

4. H. chinensis Grev. in Ann. Mag. nat. Hist. Ill, 16: 5, fig. 9 (1865). (PI. 41, Fig. 10; specimen
length 27-5 /urn, height from base to tips, 32-5 /um)

Hong Kong Harbour 9, BM 4506, syntypes.

The drawing in the Greville manuscript collection is of three specimens. One is a single frustule
without a girdle, the second is a single frustule with an expanded girdle, and the third, the one
Greville published, is of two entire frustules. There is no indication from which slide the
published illustration was taken. These specimens are considered syntypes.

5. H. crenatus Grev. in Trans, microsc. Soc. Lond. 13: 101, fig. 12 (1865).
Cambridge, Barbadoes, BM 3444, holotype.

It is impossible to photograph the type specimen as the coverslip has become completely
detached from the slide. Any attempt to remount the coverslip will certainly move the specimen
from its original MF co-ordinates.

36 D. M. WILLIAMS

6. H. exiguus Grev. in Trans, microsc. Soc. Lond. 13: 29, fig. 20 (1865). (PI. 41, Figs 1, 2;
specimen length 26-5 /am)

Barbados Johnson 1863, BM 3057; Barbadoes Johnson 1864, BM 3128, holotype?

BM 3057 has only a single valve, whereas there is an entire frustule on BM 3128. It is possible

that the latter specimen is the holotype.

7. H. hastatus Grev. in Trans, microsc. Soc. Lond. 13: 31, fig. 15 (1865). (PI. 41, Fig. 3;
specimen length 62 /am, height from base to tips, 98-5 /am)

Barbadoes 1864, BM 3036, BM 3044, lectotype; Barbadoes Johnson 1864, BM 3037; Barbadoes
Johnson 1865, BM 3449; Barbadoes 1865, BM 3460.

Fenner (1982) transferred H. hastata to Odontella but the combination is invalid as it appeared in
an unpublished Ph. D dissertation. A valid transfer appears in the appendix.

8. H. lobatus Grev. in Trans, microsc. Soc. Lond. 13: 29, fig. 9 (1865). (Not illustrated)
Barbadoes Johnson 1864, BM 3420; Cambridge Barbadoes 1865, BM 3448.

9. H. longicornis Grev. in Trans, microsc. Soc. Lond. 13: 31, fig. 13 (1865). (PI. 40, Fig. 10;
specimen length 44-5 /am, height from base to tips, 127-5 /am)

Barbadoes 1865, BM 3115, holotype? BM 3469; Cambridge Barbadoes Johnson 1863, BM 3055;
Barbadoes Johnson 1864, BM 3266, BM 3272; [Barbadoes Johnson 1864], BM 3409; Barbadoes
Cambridge estate, BM 3443; Cambridge Barbadoes 1865, BM 3444; Barbadoes Johnson 1865,
BM 3449.

There is a drawing in Greville's manuscript collection which is labelled Monogramma ventri-
cosa, but this name has since been crossed out. The drawing is identical to that published as
H. longicornis and the slide number, given as BM 3115, specimen MF 15/22, is probably the
holotype.

10. H. lyriformis Grev. in Trans, microsc. Soc. Lond. 13: 30, figs 11, 21 (1865). (PI. 41, Fig. 4;
specimen length 33 /am, height from base to tips, 55-5 /am)

Barbadoes 1865, BM 3105, lectotype, BM 3264; [Barbadoes Johnson 1864], BM 3409;
Barbadoes Johnson 1864, BM 3421; Barbadoes Johnson 1865, BM 3439, BM 3440; Cambridge
Barbadoes 1865, BM 3444.

11. H. minutus Grev. in Trans, microsc. Soc. Lond. 13: 101, fig. 5 (1865).
Cambridge Barbadoes 1865, BM 3444.

It is impossible to photograph the type specimen as the coverslip has become completely
detached from the slide. Any attempt to remount the coverslip will certainly move the specimen
away from its MF co-ordinates.

12. H. mucronatus Grev. in Trans, microsc. Soc. Lond. 13: 28, fig. 6 (1865). (PI. 41, Figs 5-7;
specimen length 52-5 /am, height from base to tips, 35 /am)

Barbadoes Johnson 1864, BM 3266, lectotype, BM 3282, BM 3429; Cambridge Barbadoes
Johnson 1865, BM 3274, BM 3412; Barbadoes Cambridge Estate Johnson, BM 3410; Barbadoes
Johnson 1865, BM 3415; Barbadoes Cambridge estate Johnson 1864, BM 3416, BM 3419;
Cambridge Barbadoes 1865, BM 3442.

13. H. ornithocephalus Grev. in Trans, microsc. Soc. Lond. 13: 32, fig. 16 (1865). (PI. 41, Figs 8,
9; specimen length 37-5 /am, height from base to tips, 15 /am)

Barbadoes 1865, BM 3263, holotype.

There is no drawing in Greville's manuscript collection. However, this is the only specimen
present in Greville's slide collection and is considered the holotype by Ross (1972: 335) and Ross
& Sims (1985: 310).

14. H. pulvinatus Grev. in Trans, microsc. Soc. Lond. 13: 28, fig. 8 (1865). (PI. 42, Figs 1, 4;
specimen length 53 /am, height from base to tips, 32-5 /am)

Barbadoes Johnson 1864, BM 3255, holotype.

15. H. punctatus Grev. in Trans, microsc. Soc. Lond. 13: 28, fig. 7 (1865).
'Barbadoes deposit, Cambridge estate; in slides communicated by C. Johnson, Esq.'

GREVILLE DIATOM TYPES 37

There is no slide in the Greville collection of this taxon. The type slide has not been traced.

16. H. reticulatus Grev. in Trans, microsc. Soc. Lond. 13: 27, fig. 5 (1865).
'Barbadoes deposit, Cambridge estate; in slides communicated by C. Johnson, Esq.'
There is no slide in the Greville collection of this taxon. The type slide has not been traced.

17. H?? robustus Grev. in Trans, microsc. Soc. Lond. 13: 54, fig. 23 (1865). (PI. 42, Fig. 2;
specimen length 38 /xm, height from base to tips, 72-5 /xm)

Barbadoes Cambridge Estate Johnson 1864, BM 3422, holotype.
This slide is badly damaged and gives extremely poor micrographs.

18. H. symmetricus Grev. in Trans, microsc. Soc. Lond. 13: 53, fig. 22 (1865). (PI. 42, Figs 6, 7;
specimen length 90 /am, height from base to tips, 30 /am)

Barbadoes Johnson 1864, BM 3282, holotype.

The drawing in the Greville collection was incorrectly annotated. The holotype specimen has
been found on the above slide. According to Mr R. Ross, careful examination of the type
specimen shows that it is a broken specimen of a tripolar Trinacria sp.

19. H. tenuicornis Grev. in Trans, microsc. Soc. Lond. 13: 29, fig. 10 (1865). (PI. 43, Figs 1, 2;
specimen length 60 /xm, height from base to tips, 80 /xm)

Barbadoes 1865, BM 3264, lectotype, BM 3280; Barbadoes, Johnson, 1864, BM 3285.
There is no drawing for this species in the Greville manuscript collection.

XL VI. HETERODICTYONGrev.

1. H. rylandsianum Grev. in Trans, microsc. Soc. Lond. 11: 66, fig. 6 (1863). (PI. 42, Figs 3, 5;
specimen diameter 102-5 /xm)

Barbadoes Johnson 1862, BM2771, holotype.

2. H. splendidum Grev. in Trans, microsc. Soc. Lond. 11: 66, fig. 7 (1863). (PI. 42, Fig. 8;
specimen diameter 42-5 /xm)

Barbadoes Johnson 1853 [sic, 1863], BM2820, holotype.

XL VII. LIRADISCUSGrev.

1. L. barbadensis Grev. in Trans, microsc. Soc. Lond. 13: 5, fig. 14 (1865). (PI. 42, Fig. 9;
specimen diameter 60 /xm)

Barbadoes Johnson 1864, BM 3179, holotype.

2. L. ellipticus Grev. in Trans, microsc. Soc. Lond. 13: 99, fig. 6 (1865). (PI. 42, Fig. 10;
specimen diameter 73 /xm)

Cambridge Barbadoes Johnson 1865, BM 3274, holotype.

Two specimens are indicated on the slide. The specimen at MF location 32/18 is indicated as

holotype on Greville's drawing.

3. L. minutus Grev. in Trans, microsc. Soc. Lond. 13: 47, fig. 6 (1865). (PI. 43, Fig. 3; specimen
length 17-5 /xm)

Barbadoes Johnson 1864, BM 3281, BM 3275, syntypes.

The published figure is of three separate drawings. In the Greville manuscript collection two
drawings remain. BM 3275 has a single specimen (MF 32/21, Greville's figure 6, lower-most
valve; illustrated) and BM 3281 has two specimens, MF 33/22 (Greville's figure 6, uppermost
valve) and MF 19/21 (Greville's figure 6, middle valve). The drawing with two valves has BM
3295 added, but clearly in error, whilst the other drawing denotes only one MF number.

4. L. ovalis Grev. in Trans, microsc. Soc. Lond. 13: 5, figs 15, 16 (1865). (PI. 43, Fig. 4;
specimen diameter 47-5 /xm)

Cambridge Barbadoes Johnson 1864, BM 3181, BM 3182, syntypes.

Greville illustrates two specimens and both figures are present in the Greville manuscript

collection. The valve view is annotated with slide numbers BM 3181 and BM 3182, whilst the

38 D. M. WILLIAMS

girdle view is without a slide reference. Each slide has several specimens. The micrographs are
from BM 3182.

XLVIII. MASTOGLOIA Thw. ex W. Sm.

1. M. capitata Grev. in Q. Jl microsc. Sci. II, 2: 235, figs 11, 12 (1862). (PI. 43, Figs 5, 6, 9;
specimen length 32-5 /am)

Tank Calcutta Dr Macrae 1861, BM 2563, BM 2564, syntypes.

There are only two slides in the collection that correspond to the type locality and specimens are

abundant on each slide.

2. M. macdonaldii Grev. in Trans, hot. Soc. Edinb. 8: 237, fig. 15 (1866).
'Shark's Bay, Australia, in stomachs of Ascidians. Dr Macdonald.'

There is a drawing in the Greville manuscript collection which indicates that the slide belonged
to Mr Norman. The drawing is annotated with: 'Shark's Bay. Norman's Slide. 810. 28/35'. This is
clearly a reference to the holotype specimen, but is has not been located in the collections at BM.

3. M. minuta Grev. in Q. Jl microsc. Sci. 5: 12, fig. 10 (1857). (PI. 43, Figs 7, 8; specimen length
28-5 /am)

Trinidad Mrs Eccles 9.55, BM 995; Trinidad 1856, BM 1002, syntypes.
Greville stated that he had examined numerous frustules.

XLIX. MELOSIRA Agardh

1. M. costata Grev. in Trans, microsc. Soc. Lond. 14: 77, figs 3-6 (1866). (PI. 44, Figs 1,2; valve
width, 11 /am, frustule length 24-5 /am)

Hong Kong, BM 4501, BM 53909, syntypes.

Hasle (1973) made an extensive study of this species. She used material from Hong Kong (BM
53909) for her study, which was also from J. Linton Palmer's original gathering. The additional
locality from Yorkshire, which Greville received from G. Norman, has not been traced.

2. M. nobilis Grev. in Trans, hot. Soc. Edinb. 8: 233, fig. 2 (1866). (PI. 43, Figs 10, 11; specimen
width 75 /am, frustule length 82-5 /am)

New Caledonia Dr Roberts, BM 2661, holotype.

Only two specimens are indicated in the collection, both of which are on the same slide. The

micrographs are from the specimen that Greville illustrated. The other specimen is of two linked

frustules.

3. M. setosa Grev. in Trans, hot. Soc. Edinb. 8: 436, figs 17-19 (1866). (PI. 44, Figs 3, 4; frustule
length 28-5 /am)

Aneitum New Hebrides, BM 3532, BM 3533, BM 3534, syntypes.

L. MONOGRAMMA Ehrenb.

1. M. smithiana Grev. in Trans, hot. Soc. Edinb. 8: 440, figs 3-5 (1866). (Not illustrated)
Aneitum New Hebrides, BM 3532, BM 3533, BM 3534, syntypes.

LI. NAVICULA Bory

1. N. approximata Grev. in Edinb. Newphil. J. 10: 28, fig. 4 (1859); in Trans, hot. Soc. Edinb. 6:

247, fig. 4 (1859). (PI. 44, Fig. 15; specimen length 82-5 /am)

Californ. Guano, BM 1765, BM 1774, lectotype, BM 1775; Californ. Guano,/. T. Norman, BM
1767; Californ. Guano,/. T. Norman 1858, BM 1773; [Californ. Guano], BM 1776.
There is a single drawing in the Greville manuscript collection which is annotated with two
numbers: 21(5) and 22 (2). Neither of these numbers are traceable on slides.

2. N. californica Grev. in Edinb. Newphil. J. 10: 29, fig. 5 (1859); in Trans, hot. Soc. Edinb. 6:

248, fig. 5 (1859). (PI. 44, Figs 13, 14; specimen length 115 /am)
So. African Guano, BM 1859, holotype?; Patos guano, BM 1766.

GREVILLE DIATOM TYPES 39

Greville states in the protologue: 'an individual from South African Guano now before me',
suggesting that the specimen on BM 1859 was used for the published illustration.

3. N? cistella Grev. in Trans, microsc. Soc. Lond. 11: 19, figs 12-14 (1863). (PI. 44, Fig. 5;
specimen length 28 /xm; PI. 44, Fig. 6; specimen length 29-5 /xm)

Harvey Bay, BM 2665, BM 2669; Harvey Bay Dr Roberts 1862, BM 2676, BM 2677, BM 2678,
BM 2679; Dredged in 8 fath. Lyme Regis, Roper, BM 2704, syntypes.

Greville illustrated this species with three drawings. Figures 13 and 14, the valve views, are
missing from the Greville manuscript collection, but figure 12, a girdle view, is present.
However, no slide information is given. Greville gives the following localities in the protologue:
'Dredged off Lyme Regis, in eight fathoms' water by the Rev. J. Guillemard, 1855; F. C. S.
Roper, Esq. Harvey Bay, Queensland, in dredging communicated by Dr. Roberts, of Sydney;
not infrequent.' Greville's description is apparently based on the Harvey Bay material. Figure
12 is likely to be that of Roper's material as in the protologue Greville states that only Roper had
seen a girdle view. However, BM 2704, of Roper's material, has only valve view specimens.

4. N. compacta Grev. in Q. Jl microsc. Sci. 5: 11, fig. 8 (1857). (PI. 44, Fig. 10; specimen length
32-5 jam)

Trinidad Mrs Eccles, 9.55, BM 995 holotype.

This is the only slide in the Greville collection that has a specimen identified by Greville.

5. N. diversa Grev. in Trans, hot. Soc. Edinb. 7: 579, fig. 14 (1863); in Edinb. New phil. J. 18:
186, fig. 14 (1863). (PI. 44, Figs 8, 9; specimen length 55 /xm)

Curteis Straits Queensland Dr Roberts 1862, BM 2640, holotype.

6. N. egyptiaca Grev. in Trans, microsc. Soc. Lond. 14: 127, figs 16, 17 (1866). (PI. 44, Figs 7,
11, 12, 16; specimen length 131 /xm)

Stomach of Holothurians Alexandria Norman 1866, BM 3000, lectotype.
A drawing in the Greville manuscript collection is annotated with 'Holothuria Alexandria'. The
drawing is of three specimens, one in valve view and two in girdle view. One of the girdle view
illustrations was not published. This specimen is given in Fig. 7. The additional slides Greville
received from W. J. Baker have not been located. The above slide is the only one in the
collection for this species and it is designated as lectotype.

7. N. excavata Grev. in Trans, microsc. Soc. Lond. 14: 130, fig. 15 (1866).
'Red Sea; cabinet of Laurence Hardman, Esq.'

There is some confusion as to the source of the type illustration. The drawing in the Greville
manuscript collection is annotated as 'H. Monterey 65' yet the locality given in the protologue is
the Red Sea. There is also a set of manuscript notes annotated for Monterey. All the slides in the
Greville collection and the Hardman slides (Deby collection) from the Red Sea have specimens
of jV. rimosa. However, two slides are purported to have specimens of N. excavata, from Santa
Monica (BM 10284) and Los Angeles (BM 10290). There are no slides from Monterey that
indicate N. excavata.

8. N. forcipata Grev. in Q. Jl microsc. Sci. 7: 83, figs 10, 11 (1859). (PI. 45, Fig. 1; specimen
length 74-5 /xm)

Creswell Donkin, BM 1522, lectotype; Arran 1857, BM 1450, BM 1473, BM 1475, BM 1557,
BM 1560; Arran, BM 2231; Californ. Guano, BM 1783, BM 1810, BM 1824; Californ. Guano
/. T. Norman 1858, BM 1797; Glenshire, BM 3295.

Greville gave several localities in the protologue for this species, including: Glenshira, Lamlash
Bay, Creswell, and Californian guano. All these are represented in the collection and have been
listed above. However, Greville also noted: The Northumberland examples are the finest which
have come to my notice. Those from the Clyde are, for the most part, much smaller - many of
them even minute - thus exhibiting a range size as extensive as in N. lyra and its allies . ' Therefore
I have chosen as lectotype the Northumberland slide.

9. N. gemmata Grev. in Edinb. New phil. J. 10: 30, fig. 7 (1859); in Trans, hot. Soc. Edinb. 6:
249, fig. 7 (1859). (PI. 45, Figs 2, 3; specimen length 120 /xm)

40 D. M. WILLIAMS

Patos guano, 8.58,7. T. Norman, EM 1755 (specimen illustrated), BM 1757, BM 1771; Calif orn.
guano, 7. T. Norman, BM 1764, BM 1794, BM 1815.

10. N. gregoriana Grev. in Q. Jlmicrosc. Sci. 5: 10, fig. 7 (1857).
Trinidad, Mrs William Eccles.

All the slides in the Greville collection with specimens of this taxon are from localities other than
the type: BM 1670 and BM 1732 from Jamaica and BM 1749 from shell scrapings, Nassau,
W. Indies. The type slide has not been located.

11. N. indica Grev. in Trans, microsc. Soc. Lond. 10: 95, fig. 13 (1862). (PI. 45, Figs 4-6;
specimen length 112-5 /am)

Ind. Oc. Macrae 1861, BM 2417, holotype.

12. N. irrorata Grev. in Edinb. New phil. J. 10: 27, pi. IV, fig. 1 (1859); in Trans, hot. Soc.
Edinb. 6: 246, fig. 1 (1859). (PI. 45, Figs 7, 8; specimen length 100 /am)

Californ. Guano, BM 1775, BM 1782, BM 1810, BM 1811 (specimen illustrated); Californ.
Guano 7. T. Norman, BM 1767, BM 1794; Californ. Guano J. T. Norman 58, BM 1806;
Californ. Guano 7. T. Norman 8.58, BM 1820, BM 1779; Californ. Guano 7. T. Norman 9.58,
BM 1768, BM 1769.

The drawing is missing from the Greville manuscript collection and there is no indication of
which slide Greville used for his drawing.

13. N. jamaicensis Grev. in Trans, microsc. Soc. Lond. 14: 126, fig. 23 (1866). (PI. 45, Figs 9, 10;
specimen length 80 /urn)

Washed from shells from Aneitum, New Hebrides, BM 1692, holotype?
Greville gives Jamaica as the type locality but the only slide listed in Greville's collection is from
Aneitum, New Hebrides. On this slide a fractured specimen is present at the MF number
Greville noted on the manuscript drawing where Aneitum has been crossed out and replaced
with Jamaica. Examination of the strewn slide has shown that the species composition is
identical to the others from Aneitum.

14. N. johnsoniana Grev. in Trans, microsc. Soc. Lond. 11: 17, fig. 8 (1863). (PI. 46, Figs 1, 2;
specimen length 92-5 /am)

Harvey Bay, Queensland, BM 2673, holotype?; N. Zealand 1859 Johnson, BM 2568.
The drawing is missing from the Greville manuscript collection. Greville gives two localities in
the protologue: New Zealand, C. Johnson and Harvey Bay, Dr Roberts. However, Greville
states in the protologue: 'I had prepared drawings of both forms; but not having room in the
plate for both, I have given the Australian . . .'

15. N. lewisiana Grev. in Trans, microsc. Soc. Lond. 11: 15, fig. 7 (1863). (PI. 46, Figs 3, 5, 6;
specimen length 182 /am)

Sierra Leone F. Kitton, R321 , BM 3898; Sierra Leone, Kitton, BM 1892 (MF 24/25, illustrated).
Greville gives several localities in the protologue but only the above slides are catalogued for this
species.

16. N. luxuriosa Grev. in Trans, microsc. Soc. Lond. 11: 18, figs 10, 11 (1863).
'Port Stephen, New South Wales, in a dredging communicated by Dr. Roberts.'

Both of the published illustrations are in the Greville manuscript collection. The card with the
original drawings is annotated 'Port Stephen 3' and has two MF locations, 21/21 and 25/12. This
slide has not been located.

17. N. lyra var. recta Grev. in Edinb. New phil. J. 10: 28, pi. IV, fig. 3 (1859); in Trans, hot. Soc.
Edinb. 6: 247, fig. 3 (1859). (PI. 46, Figs 11, 12; specimen length 170 /am)

Patos guano, 8.58, 7. T. Norman, BM 1756, holotype?

The slide label is annotated with N. lyra var. princeps, a name Greville never published. Instead,
Greville published this taxon as var. recta, of which the above slide is probably the holotype.
Additionally, Greville states that he has only seen two examples. There are two adjacent
specimens on the slide.

GREVILLE DIATOM TYPES 41

18. N. notabilis Grev. in Trans, microsc. Soc. Lond. 11: 18, fig. 9 (1863). (PI. 46, Fig. 4;
specimen length 87-5 /am)

Cook's Reef Torres Strait G. Norman (381) 1862, BM 2622, lectotype.

Four specimens are indicated on the slide label. It is impossible to establish which of these

Greville used for his illustration. The specimen at MF 13/14 has been used for the micrographs.

19. N. nummularia Grev. in Edinb. Newphil. J. 10: 29, fig. 6 (1859); in Trans, hot. Soc. Edinb.
6: 249, fig. 6 (1859). (PI. 46, Fig. 7; specimen length 32-5 /am)

Californ. guano /. T. Norman, 9.58, BM 1769, BM 1773; Californ. guano 58, BM 1798,
lectotype.

20. N. polysticta Grev. in Edinb. Newphil. J. 10: 28, fig. 12 (1859); in Trans, hot. Soc. Edinb. 6:
247, fig. 2 (1859). (PI. 46, Figs 8, 9; specimen length 50 /urn)

Californ. guano, BM 1775, holotype?

21. N. rimosa Grev. in Trans, microsc. Soc. Lond. 14: 129, fig. 25 (1866). (PI. 47, Figs 1-3;
specimen length 140 /am)

Red Sea Dredgings [L. Hardman], BM 3982, syntypes.

22. N. robertsiana Grev. in Trans, hot. Soc. Edinb. 8: 235, fig. 9 (1866). (PI. 46, Fig. 10;
specimen length 167-5 /am)

New Caledonia 1862, BM 2766, holotype.

23. N. spectalissima Grev. in Trans, microsc. Soc. Lond. 14: 84, fig. 29 (1866). (PI. 47, Figs 4, 5;
specimen length 62-5 /am)

Zanzibar Prof. H. L. Smith, 65, BM 3553, holotype.

There is no drawing of this species and only one slide in the Greville collection. This is of a partial

valve.

24. N. strangulata Grev. in Trans, microsc. Soc. Lond. 14: 126, fig. 24 (1866).
'In marine dredgings, Nassau, West Indies.'

There is a drawing in the Greville manuscript collection but it is not annotated with a slide
number. There are no slides labelled for this species. The type slides have yet to be discovered.

25. N. sulcata Grev. in Trans, hot. Soc. Edinb. 8: 235, fig. 10 (1866). (PI. 47, Figs 6, 7; specimen
length 40 /am)

New Caledonia Dr Roberts 1862, BM 2655, holotype.

26. N. zanzibarica Grev. in Trans, microsc. Soc. Lond. 14: 129, fig. 28 (1866). (PI. 48, Fig. 1;
specimen length 185 /am)

Zanzibar Prof. H. L. Smith, 1865, BM 3556.

A set of manuscript notes indicates that Greville used two slides annotated as follows: 'no. 4 H.
Smith and no. 11'. The above slide is isotype material. Neither of these numbers corresponds to
slides in Greville's collection.

LII. ODONTODISCUS Ehrenb.

1. O. barbadensis Grev. in Trans, hot. Soc. Edinb. 8: 436, fig. 16 (1866). (PI. 47, Fig. 8;

specimen diameter 55 /am)

Barbados Johnson 1866, BM 3509, holotype.

LIII. OMPHALOPELTA Ehrenb.

1. O. moronensis Grev. in Trans, microsc. Soc. Lond. 14: 122, fig. 14 (1866). (PI. 48, Figs 8, 9;

specimen diameter 57-5 /am)

Moron L. H. no. 462, BM 10371, holotype.

42 D. M. WILLIAMS

LIV. OMPHALOPSISGrev.

1. O. australis Grev. in Trans, hot. Soc. Edinb. 7: 537, figs 10, 11 (1863); in Edinb. Newphil. J.

18: 36, figs 10, 11 (1863). (PI. 48, Fig. 2; specimen diameter 52-5 /am)

[Woodlark Islands Dr Roberts], BM 2714, syntypes.

The drawing in the Greville manuscript collection is annotated with '[BM] 2714'. The slide label

has no identifications or locality given; it has been left blank. There are two specimens on the

slide, MF 28/19, which corresponds to figure 10 (not illustrated) and MF 21/24, which

corresponds to figure 11 (illustrated).

LV. PALMERIA Grev.

1. P. hardmaniana Grev. in Ann. Mag. nat. Hist. Ill, 16: 2, figs 1-4 (1866). (PI. 48, Figs 4-6;

specimen diameter 485 /am)

Hong Kong Harbour, BM 2998, BM 4721, syntypes.

Stosch (1986) has recently studied this genus and given an enlarged description of both the genus

and species. He examined Greville's type material.

LVI. PEPONIA Grev.

1. P. barbadensis Grev. in Trans, microsc. Soc. Lond. 11: 76, fig. 25 (1863). (PI. 48, Fig. 7;

specimen diameter 72-5 /am)

Bridgewater Barbadoes Johnson 1863, BM 2739, holotype.

LVII. PINNULARIA Ehrenb.

1. P. hartleyana Grev. in Trans, microsc. Soc. Lond. 13: 57, fig. 30 (1865). (PI. 48, Fig. 3;

specimen length 245 /am, width 27-5 /am)

Cavalla Liberia Rev. B. Hartley, 1864, BM 3374, syntype.

This is the only slide in the collection from the type locality. There are two specimens marked on

the slide with MF locations: 26/29 and 35/28. The latter has been used for the photograph.

LVIII. PLAGIOGRAMMA Grev.

1. P? angulatum Grev. in Trans, microsc. Soc. Lond. 14: 121, fig. 3 (1866). (PI. 49, Figs 1-3;
specimen length 95 /am)

Cambridge Barbadoes Johnson 1862, BM 2758, holotype.

2. P. atomus Grev. in Trans, hot. Soc. Edinb. 7: 536, fig. 9 (1863); in Edinb. Newphil. J. 18: 36,
fig. 9 (1863). (PI. 49, Figs 4, 5; specimen length 20-5 /am)

[Woodlark Island Dr Roberts], BM 2714, holotype.

3. P. barbadense Grev. in Trans, microsc. Soc. Lond. 14: 1, fig. 3 (1866).
Cambridge Barbadoes 1865, BM 3444, holotype.

This is a badly damaged slide making it impossible to photograph the type specimen. Several
alternative slides are available for neotype specimens: BM 3187, BM 3254, BM 3419, BM 3447.

4. P. californicum Grev. in Q. Jlmicrosc. Sci. 7: 211, figs 15-17 (1859). (PI. 49, Fig. 9; specimen
length 55 /am, PI. 49, Fig. 10, specimen length, 60 /am)

Californ. guano, BM 1790; Californ. guano Arnott, (S. 599), BM 1793; Californ. guano BM

1795; [Californ. guano], BM 1776.

It is possible to match specimens on the slides to the published illustrations. BM 1790 is a

fragment and possibly that of figure 15; either BM 1793 or BM 1795 (PI. 49, Fig. 9) could be the

specimen in figure 17; and BM 1776 is possibly that of figure 16. In any case, these are all syntype

specimens.

5. P. constrict urn Grev. in Trans, hot. Soc. Edinb. 7: 536, fig. 8 (1863); in Edinb. Newphil. J. 18:
36, fig. 8 (1863). (PI. 49, Figs 6, 7; specimen length 52-5 /am)

New Caledonia, BM 2643, holotype.

GREVILLE DIATOM TYPES 43

6. P. costatum Grev. in Trans, hot. Soc. Edinb. 7: 535, figs 5, 6 (1863); in Edinb. Newphil. J. 18:
35, figs 5, 6 (1863). (PI. 49, Fig. 8; specimen length 43-5 /urn)

New Caledonia Dr Roberts, 1862, BM 2646, syntypes.

The drawing in Greville's manuscript collection indicates only one slide and a single MF

location (25/12). This specimen is illustrated by Greville's figure 6. The slide has eight other

specimens. As the original description referred to two figures, the above must be considered

syntypes.

7. P. decussatum Grev. in Trans, microsc. Soc. Lond. 14: 1, fig. 1-2 (1866). (Not illustrated)
St. Helena Wallich, 1862, BM 2579; Zanzibar Prof. H. L. Smith 65, BM 3553; Zanzibar Prof. H.
L. Smith, 65, EM 3562.

A drawing is present in the Greville manuscript collections but there is no reference to which
specimen it was taken from. Greville states: 'The discovery of this species is due to Dr. Wallich,
in whose notes and sketches it is clearly indicated.' BM 3553 and BM 3562 are of valve views
whilst the girdle view is that of BM 2579. These are all syntypes. The micrographs have been
taken from BM 3553.

8. P. elongatum Grev. in Trans, microsc. Soc. Lond. 14: 121, figs 1, 2 (1866). (PI. 49, Fig. 11;
specimen length 167-5 /am)

From shell cleanings So. America L. Hardman, 1866, BM 3490; Shell cleanings, So. America,

BM3491.

There is a drawing in the Greville manuscript collection but without annotations. BM 3490 is a

selected slide with one valve view specimen and BM 3491 is a group of selected valves with

a portion of P. elongatum in girdle view. Additionally, there is a label on 3490 which says

'Dr Greville, from L. Hardman!'

9. P. inaequale Grev. in Q. Jl microsc. Sci. 7: 210, fig. 10 (1859). (PI. 50, Figs 1-3; specimen
length 40 /urn)

Jamaica, BM 1712; Nassau, 59, BM 1736.

Greville states that the valve view is unknown. Two localities are given in the protologue, hence

these slides are syntypes. The micrographs have been taken from BM 1736.

10. P. jamaicense Grev. in Q. Jl microsc. Sci. 7: 208, fig. 3 (1859). (PI. 49, Figs 12, 13; specimen
length 38 /xm)

Jamaica 58, BM 1705, BM 1706, syntypes.

The micrographs have been taken from BM 1705.

11. P. lyratum Grev. in Q. Jl microsc. Sci. 7: 211, fig. 14 (1859). (PI. 50, Figs 4, 5; specimen
length 67-5 /xm)

Shell scrapings Nassau W. Ind. Norman, BM 1730, holotype.

Greville states: 'I have only met with one example', hence this is probably the holotype. The

specimen is only a fragment and the slide is such that only low magnification is possible.

12. P. obesum Grev. in Q. Jl microsc. Sci. 7: 211, figs 12, 13 (1859). (PI. 50, Fig. 10; specimen
length 45 /xm)

Shell scrapings Nassau Norman, BM 1731, lectotype; Nassau, BM 1737; Nassau W. Ind., BM
1740, BM 1744.

13. P. orientale Grev. in Trans, microsc. Soc. Lond. 14: 77, fig. 1 (1866). (PI. 50, Figs 8, 11;
specimen length 25-5 /xm)

Zanzibar Prof. H. L. Smith 65, BM 3553, holotype.

There is a drawing in the Greville manuscript collection but without annotations. The species is
referred to as P. africana in Greville's manuscript notes. The notes indicate which slide Greville
used for the illustration.

14. P. ornatum Grev. in Q. Jl microsc. Sci. 7: 209, fig. 9 (1859). (PI. 50, Fig. 9; specimen length
112-5 /urn)

Californ. Guano Arnott (S. 599), BM 1760, holotype?

44 D. M. WILLIAMS

15. P. pulchellum Grev. in Q. Jlmicrosc. Sci. 7: 209, figs 4-6 (1859). (PI. 50, Fig. 6; specimen
length 55 /xm)

Californ. guano 58 Norman, BM 1759; Californ. guano, BM 1785, BM 1787; Patos guano 58,

BM 1758, syntypes.

Greville published three drawings for this species, noting that he has 'seen as many as five

frustules united in a chain'. BM 1759 is figure 5; BM 1785 is figure 6; and BM 1787 is figure 4

(illustrated).

16. P. pygmaeum Grev. in Q. Jlmicrosc. Sci. 7: 211, fig. 11 (1859). (PI. 51, Figs 1, 6; specimen
length 25 /u,m)

Shell scrapings Nassau, BM 1735 holotype?

17. P. robertsianum Grev. in Trans, microsc. Soc. Lond. 11: 13, figs 1, 2 (1863).
'Port Stephen, New South Wales; Dr. Roberts.'

There are neither manuscript notes nor slides of this species in the Greville collection.

18. P. spectabile Grev. in Edinb. Newphil. J. 18: 35, fig. 7 (1863); in Trans, hot. Soc. Edinb. 7:
536, fig. 7, (1863). (PI. 51, Fig. 2; specimen length 48 Aim)

New Caledonia Dr Roberts, 1862, BM 2645, BM 2652.

Greville notes that 'of this diatom I have only seen two examples'. In the Greville manuscript
collection there is a drawing with the two numbers. However, there appear to be several
specimens on each slide. The drawing indicates BM 2652 (MF 32/22), but this specimen is only a
fragment. Additionally, BM 2645 (MF 33/35) is indicated, which corresponds to an entire valve.
The latter specimen has been used for the micrographs.

19. P? tessellatum Grev. in Q. Jl microsc. Sci. 7: 208, fig. 7 (1859). (PI. 50, Fig. 7; specimen
length 80 /u-m)

Californ. Guano, BM 1789, holotype.

20. P. validum Grev. in Q. Jlmicrosc. Sci. 7: 209, fig. 8 (1859). (PI. 51, Figs 3, 4; specimen length
110-5 /urn)

Californ. guano/. T. Norman, BM 1794, lectotype; Californ. guano/. T. Norman 58, BM 1825;
Californ. guano Am. s. 599, BM 1826.

Greville states in the protologue: 'I have only seen the side view [valve view] of this diatom'. BM
1794 has a specimen in valve view and is therefore probably the holotype.

21. P. wallichianum Grev. in Trans, microsc. Soc. Lond. 13: 1, figs 7, 8 (1865). (PI. 51, Fig. 5;
specimen length 26 /u,m; PI. 51, Fig. 7; specimen length, 37-5 /urn)

St. Helena Waltich, (7), 1857, BM 3228; St. Helena, 1857, Wallich, (9), BM 3955; St. Helena,

Wa///c/i(14),BM3939.

The drawings in the Greville manuscript collection are of the published figures 7 and 8. Figure 7

is annotated with 'no. 7, 15/29', corresponding to BM 3228, with an additional annotation to an

alternative valve view, 'St. Hel. 14, 24/32'. Figure 8 is annotated 'n.9, 20/21', which is the

specimen on BM 3955.

LIX. PLEUROSIGMA W. Sm.

1. P. compactum Grev. in Q. Jlmicrosc. Sci. 5: 12, fig. 9 (1857). (PI. 51, Figs 8-10; specimen

length 117-5 /am)

Trinidad Mrs Eccles, BM 993, holotype.

In a note Cox (1983: 579) made concerning the species, she suggests affinities with either

Rhoicosigma Grun. or Donkinia Ralfs. As authentic material has now been discovered, this can

be clarified.

LX. PORODISCUSGrev.

1. P. concinus Grev. in Trans, microsc. Soc. Lond. 11: 65, fig. 3 (1863). (PI. 52, Figs 1-3;
specimen diameter 41 /urn, frustule length 77-5 ju,m)
Cambridge Barbadoes/o/mson 1862, BM 2812, holotype.

GREVILLE DIATOM TYPES 45

2. P. elegans Grev. in Trans, microsc. Soc. Lond. 11: 64, fig. 1 (1863). (PI. 52, Fig. 4; specimen
diameter 40 /am)

Bridgewater Barbadoes Johnson, BM 2745, holotype.

3. P. major Grev. in Trans, microsc. Soc. Lond. 11: 64, fig. 2 (1863). (PI. 52, Figs 5, 6; specimen
diameter 95 /am, central area, 17-5 /am)

Cambridge Barbadoes 1863, BM 2844, holotype?

The description of this species was later amended by Greville (1865c: 46).

4. P. nitidus Grev. in Trans, microsc. Soc. Lond. 11: 65, fig. 4 (1863). (PI. 52, Figs7, 8; specimen
diameter 45 /am)

Barbadoes Johnson, BM 2751, holotype?; Cambridge Barbadoes Johnson 1863, BM 2744.
The drawing in the Greville manuscript collection is annotated with respect to two slides as the
source of the holotype: BM 2751 (MF 24/32) and BM 2744 (MF 26/31). From an examination of
these two specimens it is more likely that BM 2751 was used for the published illustration
because the specimen on BM 2744 is a partial valve.

5. P. oblongus Grev. in Trans, microsc. Soc. Lond. 11: 65, fig. 5 (1863). (PI. 53, Figs 2, 3, 5;
specimen diameter 55 /urn)

Barbadoes earth, BM 2045, holotype?

This species is referred to as P. ovalis on both the slide label and the published plate legend.

6. P. splendidus Grev. in Trans, microsc. Soc. Lond. 13: 46, fig. 5 (1865). (PI. 52, Fig. 9;
specimen diameter 35 /am)

Barbadoes deposit, Springfield Estate, L. H. no. 505, BM 10399; Barbadoes, (L. H. no. 714),
BM 10602, lectotype, (L. H. no. 715), BM 10603, (L. H. no. 716), BM 10604, (L. H. no. 717),
BM 10605.

Some of Hardman's slides are labelled confirming Greville's identifications. BM 10603 is
marked 'Greville no. 10', BM 10604 as 'Named by Greville no. 19', and BM 10605 as 'Named by
Greville no. 17'. The drawing of this species in the Greville manuscript collection is marked 'H. 3
Barbadoes' and does not relate to any of the above slides Greville identified for Hardman.

LXI. PORPEIA J. W Bail, ex Ralfs

1. P. ornata Grev. in Trans, microsc. Soc. Lond. 13: 53, fig. 21 (1865). (PI. 51, Figs 11, 12;
specimen length 73-5 /am)

Barbadoes Johnson 1864, BM 3285, holotype.

2. P. quadrata Grev. in Trans, microsc. Soc. Lond. 13: 53, fig. 20 (1865). (PI. 53, Fig. 1;
specimen length 85 /am)

Cambridge Barbadoes Johnson 1864, BM 3124, holotype.

LXH. PYXILLA Grev.

The separation of Pyrgupyxis Hendey from Pyxilla Grev., on the basis of the presence or
absence of the attachment scars, is now open to some dispute and the relationship Pyxilla has
with other similar species is still under investigation (see Hendey, 1969; Simonsen, 1979: 22;
Gombos, 1983 and Gombos & Ciesielski, 1983).

1. P. barbadensis Grev. in Trans, microsc. Soc. Lond. 13: 2, fig. 5 (1865). (PI. 53, Fig. 7;

specimen length 61 /am)

Barbadoes 1865, BM 3108, holotype.

There are two slides in Greville's collection and a drawing of the published figure in the Greville

manuscript collection which refer to this species. The drawing indicates that BM 3108 was used

for the published figure. Hendey (1969) erroneously used the alternative slide, BM 3037.

However, both are from the type locality, and BM 3037 is a paratype specimen. Simonsen (1979:

22) used BM 3037 for his discussion of this genus.

46 D. M. WILLIAMS

2. P. johnsoniana Grev. in Trans, microsc. Soc. Lond. 13: 2, fig. 6 (1865). (PI. 53, Fig. 6;

specimen length 52-5 /urn)

Barbadoes Johnson 1862, BM 2771, holotype.

LXIII. RHIZOSOLENIA Ehrenb.

1. R. striata Grev. in Trans, hot. Soc. Edinb. 8: 234, fig. 4 (1865).

Shark's Bay, west coast of Australia, in stomachs of Ascidians, Dr MacDonald, Norman 808,

(slide not found).

The drawing in the Greville manuscript collection is annotated '808 Norman'. This slide has not

been located. Sundstrom has designated BM 1948 as lectotype (Sundstrom, 1986: 84, fig. 209).

LXIV. RUTILARIA Grev.

A brief description of the genus has been given in Ross (1976) . His study is partly based on SEM
observations. A monographic revision is underway (Ross, in prep.).

1. R. elliptica Grev. in Q. Jl microsc. Sci. II, 3: 229, fig. 3 (1863). (PI. 53, Fig. 4; specimen length

Barbadoes Johnson 1862, BM 2756, holotype.

Greville illustrated this specimen in a further publication (1866d: figs 9, 10). These were

produced from specimens on BM 2789 (MF 11/29; figure 9) and BM 3434 (MF 15/20; figure 10).

2. R. epsilon Grev. in Q. Jl microsc. Sci. II, 3: 228, fig. 1 (1863). (PI. 53, Fig. 10; specimen length
162-5 /Am)

Monterey Earth Johnson, 1864, BM 3376.

Although there is a drawing in the Greville manuscript collection, there is no indication which
slide was used for the description and figure. This slide cannot be traced nor can any others
mounted before the name was published. Mr R. Ross has suggested that BM 3376, from the
original locality, could be chosen as a suitable neotype.

3. R. obesa Grev. ex Cleve in K. svenska VetensAkad. Handl. 18 (5): 19 (1881). (PI. 53, Figs 8, 9;
specimen length 150 ^im)

Monterey L. H. no. 514, BM 10407, holotype.

In the Greville manuscript collection the drawing of this species refers to a Hardman slide, 'H.

Monterey'. This slide has been identified as BM 10407 by Mr R. Ross.

4. R. superba Grev. in Trans, microsc. Soc. Lond. 14: 125, figs 11, 12 (1866). (PI. 53, Fig. 11;
specimen length 118 /urn)

Barbadoes Johnson, 1864, BM 3427, BM 2761.

The originals of Greville's two figures are both syntypes. Both drawings are extant, with BM

2761 of figure 11 and BM 3427 of figure 12.

5. R. ventricosa Grev. in Q. Jl microsc. Sci. II, 3: 228, fig. 2 (1863).

Barbadoes Johnson 1862, BM 2765; Cambridge Barbadoes Johnson 1862, BM 2775, BM 2799,
BM 2801; Barbadoes, BM 66928.

The drawing in the Greville manuscript collection is annotated with 'Ralfs'. The only relevant
slide in Ralfs' collection is BM 66928. This may be the holotype.

LXV. RYLANDSIA Grev.

1. R. biradiata Grev. in Trans, microsc. Soc. Lond. 9: 68, fig. 1 (1861). (PI. 54, Fig. 1; specimen

diameter 53 /^m)

Barbadoes deposit, BM 2064, BM 2073, BM 2074, BM 2077, BM 2108, BM 2109; Barbadoes,

BM 2105; Barbadoes deposit Ralfs, BM 2145, lectotype.

There is a drawing in the Greville manuscript collection but no indication to the slide used in the

description and figure. As the protologue states that the material came from Ralfs, BM 2145 has

been designated lectotype.

GREVILLE DIATOM TYPES 47

LXVI. SKELETONEMA Grev.

1. S. barbadense Grev. in Trans, microsc. Soc. Lond. 13: 43, fig. 1 (1865). (PI. 54, Figs 8, 9;
valve diameter 25 /xm, specimen length, 82-5 /u,m)
Barbadoes Johnson 1864, BM 3059, holotype.

LXVII. SPATANGIDIUMBreb.

1. S. ralfsianum G. Norman ex Grev. in Q. Jl microsc. Sci. 7: 161, figs 7, 8 (1859). (PI. 55, Figs

1-3; specimen diameter 82-5 /zm)

Californ. guano 58, BM 1798, BM 1823, BM 1824; Californ. guano, Norman, 58, BM 1759,

lectotype; Californ. Guano,/. T. Norman, 9.58, BM 1769, BM 1773; Bolivian guano, BM 1721,

BM 1726.

The protologue gives two localities for this species, both of which are represented above.

LXVIII. STAURONEISEhrenb.

1. S. apiculata Grev. in Edinb. New phil. J. 10: 30, fig. 8 (1859); in Trans, hot. Soc. Edinb. 6:
249, fig. 8 (1860). (PI. 54, Figs 2, 3; specimen length 45 /xm)

Patos guano, 8.58, BM 1753.

There is a drawing in the Greville manuscript collection but no indication of which slide was used
for the description and figure. This is the only slide in Greville's collection that corresponds to
the locality data in the protologue.

2. S. australis Grev. in Trans, hot. Soc. Edinb. 7: 579, fig. 13 (1863); in Edinb. New phil. J. 18:
187, fig. 13 (1863). (PI. 54, Fig. 7; specimen length 110 /*m)

Harvey Bay Dr. Roberts 1862, BM 2678, holotype.

3. S. decora Grev. in Trans, hot. Soc. Edinb. 8: 236, fig. 11 (1866). (PI. 54, Fig. 4; specimen
length 118-5 /xm)

New Caledonia Dr Roberts 1862, BM 2663, holotype.

4. S(?) obesa Grev. in Trans, hot. Soc. Edinb. 8: 237, fig. 12 (1866). (PI. 54, Figs 5, 6; specimen
length 43 /u,m)

[Curteis Straits Queensland Dr Roberts], BM 2736, holotype.

5. S. rotundata Grev. in Trans, microsc. Soc. Lond. 14: 85, figs 30, 31 (1866). (PI. 55, Fig. 4;
specimen length 73 jam)

New Zealand [Dr Lauder Lindsay], BM 4129.

The drawing in the Greville manuscript collection is annotated with 'no. 8'. Greville had made a
series of slides from the New Zealand material with most not fully labelled. BM 4129 is no. 8 of
the series and the specimens therein are considered syntypes.
This species was published by Lauder (1865: 134), prior to Greville, as a nom. nud.

6. S. scaphulaeformis Grev. in Trans, microsc. Soc. Lond. 14: 86, fig. 32 (1866). (Not
illustrated)

Otago New Zeal. Dr Lindsay, 1861 (15), BM 3549, holotype.

The drawing in the Greville manuscript collection is annotated 'no. 15, 13/25'. This corresponds
with BM 3549 which is the holotype. However, no specimen was located at MF 13/25.
This species was published by Lauder (1865: 134), prior to Greville, as a nom. nud.

LXIX. STICTODESMIS Grev.

1. S. australis Grev. in Trans, hot. Soc. Edinb. 1: 535, figs 1-4 (1863). (PI. 55, Figs 5-7;
specimen length 220 /urn)

Harvey Bay Queensland Dr Roberts, 1863, BM 2642, lectotype, designated by Cox (1982: 149).
In a recent review of the genus Climaconeis Grun., Cox (1982) examined the type slides of
Stictodesmis australis and designated BM 2642 as lectotype. However, she did not publish

48 D. M. WILLIAMS

illustrations from this slide. Her illustrations were a drawing of the specimen from BM 2665 and
a LM from BM 3402.

LXX. STICTODISCUSGrev.

1. S. buryanus Grev. in Trans, microsc. Soc. Lond. 9: 40, fig. 1 (1861). (PI. 55, Fig. 9, PI. 56,
Fig. 1; specimen diameter 90 ^tm)

Trinidad deposit Mrs Bury, BM 2249, holotype.

2. S. californicus Grev. in Trans, microsc. Soc. Lond. 9: 79, fig. 1 (1861). (PI. 55, Fig. 8, PI. 56,
Fig. 2; specimen diameter 85 /am)

Monterey stone, BM 1971, holotype.

3. S. hardmanianus Grev. in Trans, microsc. Soc. Lond. 13: 98, fig. 4 (1865). (PI. 56, Fig. 4;
specimen diameter 117 /xm)

Monterey; L. H. no. 279, BM 10221, holotype?

The drawing in the Greville manuscript collection is annotated 'H. 49'. For further notes on this

slide see under Coscinodiscus elegans.

4. S. insignis Grev. in Trans, microsc. Soc. Lond. 9: 41, fig. 4 (1861). (PI. 55, Fig. 10; specimen
diameter 35 /xm)

Barbadoes deposit, BM 2075, holotype.

The slide is made of thick glass and hence high magnification is impossible to use and detail is

lost.

5. S. johnsonianus Grev. in Trans, microsc. Soc. Lond. 9: 41, fig. 3 (1861). (PI. 56, Fig. 3;
specimen diameter 83 /tm)

Trinidad deposit Mrs Bury, BM 2248, holotype.

The slide is made of thick glass and hence high magnification is impossible to use and detail is

lost.

6. S. kittonianus Grev. in Trans, microsc. Soc. Lond. 9: 79, figs 2, 3 (1861). (Not illustrated)
Nottingham deposit U.S., BM 2195; Richmond deposit U.S., Kitton, BM 2254, syntypes.

In the Greville manuscript drawings, a Kitton slide is indicated for the girdle view, fig. 3. This
matches the specimen on slide BM 2254. Figure 2 is the specimen on BM 2195 (MF 30/35); the
drawing for figure 3 is not annotated.

LXXI. STRANGULONEMA Grev.

1. S. barbadense Grev. in Trans, microsc. Soc. Lond. 13: 44, fig. 2 (1865). (PI. 56, Figs 5, 6;
valve width 20 /xm, specimen length 100 /xm)
Barbadoes Johnson 1863, BM 2099, holotype.

LXXII. SURIRELLA Turpin
1. S. eximia Grev. in Q. Jl microsc. Sci. 5: 10, fig. 6 (1857). (PI. 56, Fig. 8; specimen length 95

Trinidad Mrs Eccles, BM 998, lectotype.

There is no drawing of this species in the Greville manuscript collection. On the slide indicated
above two specimens are noted. The micrograph is the specimen most like Greville's drawing.
The other specimen has a distinct central constriction which, according to Greville, is part of the
variation pattern of this species.

2. S. macraeana Grev. in Trans, microsc. Soc. Lond. 10: 20, fig. 1 (1862). (PI. 56, Fig. 7;
specimen length 190 />im)

Ceylon Dr Macrae, BM 2384, holotype.

3. S. palmeriana Grev. in Ann. Mag. nat. Hist. Ill, 16: 4, fig. 7 (1865). (PI. 56, Fig. 9; specimen
length 130 /am)

Hong Kong L.H. no. 876, BM 10753, holotype.

GREVILLE DIATOM TYPES 49

4. S. superba Grev. in Trans, hot. Soc. Edinb. 8: 436, figs 1-2 (1866). (PI. 57, Fig. 1; specimen
length 142 /xm)

Cuba [H. L. Smith], BM 3530, BM 3531, BM 4670, BM 4689.

Greville states in his protologue that this species is similar to Surirella smithii Ralfs; BM 4670 and
BM 4689 are annotated as S. superba. However, these slides are prepared from the same
material that was used to make BM 3530 and BM 3531 which are labelled 5. smithii. All the
specimens Greville refers to S. superba on BM 4670 and BM 4689 and those he refers to as

5. smithii on BM 3530 and BM 3531 should be considered as syntypes.

LXXIII. SYNDETOCYSTIS Ralfs ex Grev.

This genus, discovered and named by Ralfs, was described by Greville (1866d: 125), but he did
not name the species on which it is based. This description is accurate, but Greville had earlier
sent to a correspondent in America a brief and inadequate description accompanied by an
inaccurate figure. These were reproduced by Walker & Chase (1887), who gave it a specific
name.

1. S. grevilleanus Walker & Chase, Some new and rare diatoms II-III: 6, pi. 4, fig. 13 (1887)

(S. barbadensis Ralfs ex Van Heurck, Treat. Diat. : 432, fig. 155 (1896).)

Cambridge Estate, Barbadoes, BM 2775, BM 2791, BM 2795, BM 2797, BM 2850, BM 2842,

BM2863,BM2871,BM2899,BM2902,BM2981,BM3036,BM3050,BM3061,BM3111,BM

3167, BM 3203, BM 3213, BM 3250, BM 3552, BM 3254, BM 3267, BM 3268, BM 3281, BM

3432, BM 3434, BM 3443, BM 3446, BM 3511.

The holotype of this species is clearly the original of the figure reproduced by Walker & Chase.

This was on a slide belonging to G. Norman, which cannot be traced.

LXXIV. SYNEDRA Ehrenb.

1. S. clavata Grev. in Trans, microsc. Soc. Land. 13: 25, fig. 4 (1865). (PI. 57, Fig. 2; specimen
length 185 /urn)

Barbadoes Cambridge Estate Johnson, BM 3410, BM 3441, BM 3469.

Greville states that 'not more than 4 or 5 specimens having occurred in the hundreds of slides I

have examined.' The slides above are the only ones that are noted by Greville and are probably

syntypes.

2. S. normaniana Grev. in Q. Jl microsc. Sci. II, 2: 231, figs 1-3 (1862). (PI. 57, Figs 3-5;
specimen length 175 /am)

Honduras G. Shadbolt?, 1862, BM 2543; Honduras, Smith [presented to me by Mr Roper, May
1862], BM 2542.

A detailed study of this species has been recently undertaken by Wahrer, Fryxell & Cox (1985).
They have transferred the species to Licmophora Agardh.

LXXV. SYRINGIDIUM Ehrenb.

A detailed study of Syringidium has been undertaken by Ross & Sims (1980) and Hasle & Sims
(1985). 5. daemon Grev. is considered to be the resting spore of Cerataulina bicornis (Ehrenb.)
Hasle (C. daemon (Grev.) Hasle).

1. S. daemon Grev. in Trans, microsc. Soc. Lond. 14: 83, fig. 22-28 (1866).

Hong Kong Harbour Dr Palmer, 1864, BM 4500; Hong Kong L.H. no. 567, BM 10452,

syntypes.

The drawing in the Greville manuscript collections has all seven specimens, each with a MF

location number. All specimens are from BM 4500.

Several of Greville's specimens have been examined and published: in Ross & Sims (1980: pi.
1, fig. 5) a specimen from BM 4500, and in Hasle & Sims (1985: fig. 3) a specimen from BM 52587
(see also figs 16 and 17 for material from Hong Kong Harbour, Aug. (1863) ex Lauder).

Hasle & Syvertsen (1980) illustrate a different specimen from BM 52587 (1980: pi. 5, fig. 30);

50 D. M. WILLIAMS

see also plates 7-9 for details of the vegetative valves of this species. The material from BM
52587 is from Greville's original gathering.

LXXVI. TEREBRARIA Grev.

1. T. barbadensis Grev. in Trans, microsc. Soc. Land. 12: 9, figs 12, 13 (1864). (PI. 57, Figs 6, 7;

specimen length 130 /am)

Cambridge Estate Barbadoes Johnson, BM 2799, syntype, BM 3444; Barbadoes, Johnson,

1864, BM 3188; Barbadoes, 1865, BM 3264, BM 3276.

There are two drawings in the Greville manuscript collection, each corresponding to a published

figure. Only figure 12 has been annotated, with '[BM] 2799, [MF] 31/22'. The specimens on the

slides are listed as girdle views. The remaining slide, BM 3444, is badly damaged and cannot be

inspected. This taxon is probably a Polycistine (Ross, ms notes).

LXXVII. THAUMATONEMA Grev.
This genus has recently been reviewed by Ross & Sims (1985).

1. T. barbadense Grev. in Trans, microsc. Soc. Lond. 11: 76, fig. 26 (1863). (PI. 57, Fig. 9;
specimen length 84 /u,m)

Cambridge Barbadoes Johnson 1863, BM 2853, holotype.

2. T? costatum Grev. in Trans, microsc. Soc. Lond. 13: 97, fig. 3 (1865). (PI. 57, Fig. 8;
specimen diameter 48 /xm)

Barbadoes Cambridge Estate Johnson 1864, BM 3419, holotype.

LXXVIII. TRICERATIUMEhrenb.

This genus is clearly a heterogenous mixture of unrelated taxa and in need of revision. Recent
work by Gleser (1975, 1986) proposed a revised classification into five genera: Pseudotriceratium
Grun., Trigonium Cleve, Amphitetras Ehrenb., a redefined concept of Triceratium, arid a new
genus Sheshukovia Gleser. Additionally, Jouse proposed a new genus Lisitzinia for the smaller,
cruciform species (Jouse, 1978). Many of Greville's Triceratium spp. are now classified in the
genus Entogonia Ehrenb. following the revisions by Bergon (1892a, b) and Holmes & Brigger
(1979). Further work on some species included in Triceratium by Greville will shortly be
published by Ross & Sims (in prep.).

Greville's species cut across all of these newly defined genera. No attempt has been made to
effect any taxonomic transfers of his species, except where the combinations have been made
prior to this study. References have been included where relevant.

All measurements are taken along an edge of the valve except where otherwise stated.

1. T. abercrombieanum Grev. in Trans, microsc. Soc. Lond. 9: 83, figs 7-9 (1861). (PI. 58, Figs
1,2; length 52 -5 /Am)

Barbadoes Earth, BM 2042, lectotype designated by Bergon (18926: 133).
There are three illustrations with the published description and three drawings present in the
Greville manuscript collection. Holmes & Brigger (1979) discuss Bergon's lectotypfication in
detail. Two other pertinent slides are BM 2096 and BM 2102.

Bergon (18926) referred Greville's figure 7 to T. abercrombianum, believing the other
specimens, figures 8 and 9, to be different taxa. The specimen in figure 7 is found on BM 2042
and is the lectotype designated by Bergon (1892: 133) . Holmes & Brigger (1979) accept Bergon's
lectotypification but do not concur with his concept of the species; they are of the opinion that
figure 9 (BM 2096) should be included, and have expanded the description.

2. T. acceptum Grev. in Trans, microsc. Soc. Lond. 13: 9, fig. 21 (1865). (PI. 58, Fig. 3; length 53
tun)

Cambridge Barbadoes Johnson 1863, BM 3082, holotype.

3. T. aculeatum Grev. in Trans, microsc. Soc. Lond. 9: 45 (1861). (PI. 58, Fig. 4; length 57-5
/mi)

GREVILLE DIATOM TYPES 51

Barbadoes deposit, BM 2092, holotype.

This species was described but not illustrated by Greville. The drawing Greville intended to use
is in the Greville manuscript collection. The specimen illustrated here has been identified as the
holotype by Mr R. Ross.

Ehrenberg (1856) used the combination Triceratium aculeatum in 1856, applying it to a
different taxon. In 1959 Shibkova applied it to yet another taxon (in Krotov & Shibkova, 1959).
Greville's species should be cited as T. aculeatum Grev., non Ehrenb. (1856), nee Shibkova in
Krotov & Shibkova (1959), and a new specific epithet is required.

4. T. acutangulum Grev. in Trans, microsc. Soc. Lond. 12: 93, fig. 12 (1864), non Strelnikova
(1974). (PI. 58, Figs 5, 8; length 76 /am)

Cambridge Barbadoes Johnson 1864, BM 3139, holotype.

This species belongs to Trinacria. According to Ross & Sims (in prep.) it is a later synonym of

Triceratium cornutum Grev.

5. T. amoenum Grev. in Trans, microsc. Soc. Lond. 9: 75, fig. 7 (1861). (PI. 58, Figs 9, 10; length
47-5 tun)

Nottm. deposit Maryland, BM 2163, holotype.

There are six specimens marked on the slide. The micrograph is the specimen indicated on the

drawing (MF 18/13).

6. T. approximatum Grev. in Trans, microsc. Soc. Lond. 9: 84, fig. 11 (1861). (PI. 58, Fig. 11;
length 66 /Am)

Barbadoes deposit Johnson, BM 2103, holotype.
See Holmes & Brigger (1979: 176-179).

7. T. araneosum Grev. in Trans, microsc. Soc. Lond. 13: 33, fig. 17 (1865). (PI. 59, Fig. 1; length
35 /Am)

Barbadoes 1865, BM 3262, holotype.

8. T. areolatum Grev. in Trans, microsc. Soc. Lond. 9: 71, fig. 13 (1861). (PI. 59, Fig. 2; length
62-5 (Jim)

Barbadoes deposit, BM 2087, holotype.

9. T. atomus Grev. in Trans, microsc. Soc. Lond. 13: 10, fig. 22 (1865). (PI. 59, Fig. 3; length 8
/Am)

Barbadoes Johnson 1864, BM 3200, holotype.

10. T. attenuatum Grev. in Trans, microsc. Soc. Lond. 12: 91, fig. 10 (1864). (PI. 58, Figs 6, 7;
length 60 /AITI)

Barbadoes 1864, BM 3042, holotype.

11. T. barbadense Grev. in Trans, microsc. Soc. Lond. 9: 44, fig. 12 (1861). (Not illustrated)
Barbadoes deposit, BM 2049, BM 2079.

12. T. blanditum Grev. in Trans, microsc. Soc. Lond. 9: 45 (1861); in Trans, microsc. Soc.
Lond. 9: 72, fig. 17, (1861). (PI. 59, Fig. 4; distance across centre of valve 30 /Am)
Barbadoes deposit, BM 2080, holotype.

13. T. brevinervum Grev. in Trans, microsc. Soc. Lond. 13: 101, fig. 26 (1865). (PI. 59, Fig. 5;
length 45 /Am)

Cambridge Barbadoes Johnson 1864, BM 3183, holotype.

Greville named this species 'Triceratium verrulosum var. ?' on the slide label.

14. T. browneanum Grev. in Trans, microsc. Soc. Lond. 9: 72, fig. 16 (1861). (PI. 59, Fig. 6;
length 42-5 /Am)

Mud Savannah Georgia Brown 60, BM 2161, holotype.

The manuscript drawing and the slide label refer to six specimens. MF 28/21 is identified by

Greville as the specimen used for the published illustration.

52 D. M. WILLIAMS

15. T. cancellatum Grev. in Trans, microsc. Soc. Lond. 13: 9, fig. 17 (1865). (PI. 59, Fig. 7:
length 87-5 /urn)

Barbadoes 1865, BM 3111, holotype.

The specimen occurs at the edge of the coverslip, which is cracked and the mountant receding,
and is difficult to photograph. This species should correctly be placed in Trinacria, and is being
studied by Ross & Sims (in prep.).

16. T. capitatum Ralfs ex Grev. in Trans, microsc. Soc. Lond. 9: 43, fig. 10 (1861). (PI. 59, Figs
8, 9; distance between the lower ocelli, 100 /urn)

Barbadoes deposit, BM 2024, BM 2085, BM 2087, BM 2097; [Barbadoes], 624, BM 49079,
holotype?

On the legend beneath the plate Greville has stated: The Barbadoes species are described from
a fine series of slides supplied by Mr. J. T. Norman.' The text states that 'he had examined half a
dozen examples before I perceived any trace of puncta at all. Mr. Rylands then kindly
communicated a specimen.' A slide in the Rylands collection, BM 49079, has a specimen of
T. capitatum at MF 38/24. Greville states that he found two further specimens. The micrographs
are from BM 2024.

17. T. cellulosum Grev. in Trans, microsc. Soc. Lond. 9: 44, fig. 14 (1861). (Not illustrated)
Barbadoes deposit, BM 2075, holotype.

The specimen is tilted at such an angle as to prevent useful micrographs being taken.

18. T. cinnamomeum Grev. in Q. Jl microsc. Sci. II, 3: 232, fig. 12 (1863). (PI. 59, Figs 10, 11;
length 67-5 /xm)

Moron deposit 1863, BM 2908, holotype.

A note on the original drawing indicates that specimen MF 14/19 has been used for the
illustration. One other specimen is present on this slide (PI. 60, Figs 1, 2, length, 42-5 /am). See
Simonsen (1974: 23; 1979: 66.)

19. T. concinnum Grev. in Trans, microsc. Soc. Lond. 12: 13, fig. 7 (1864). (PI. 59, Fig. 12;
length 62-5 /xm)

Cambridge Barbadoes Johnson 1862, BM 2803, holotype.

20. T. constans Grev. in Q. Jl microsc. Sic. II, 3: 233, fig. 17 (1863). (PI. 60, Fig. 3; length
77-5 /Ltm)

Cambridge Estate Barbadoes Johnson 1862, BM 2800, BM 2801; Barbadoes deposit Johnson,
BM 2804, BM 2824, BM 2849; Barbadoes deposit 1863, BM 3029; Cambridge Barbadoes
Johnson 1863, BM 3046; Barbadoes Johnson 1863, BM 3054, BM 67374; Cambridge
Barbadoes, 1863, BM 3070, syntypes.

The drawing in the Greville manuscript collection is annotated 'no. 2' and 'from Mr. Ralfs' slide
Barbadoes (Cambridge)'. There is only one slide in Ralfs' collection (BM 67374) that has
specimens of this species. The slide was originally Greville's and may be referable to the
annotations of the manuscript drawing. The specimen is a partial valve. The micrographs in this
work are taken from BM 2801.

21. T. convexum Grev. in Trans, microsc. Soc. Lond. 10: 27, fig. 6 (1862). (Micrograph not
included)

Ceylon Dr Macrae, BM 2386, holotype.

22. T. cornutum Grev. in Trans, microsc. Soc. Lond. 9: 45 (1861); in Trans, microsc. Soc.
Lond. 9: 69, fig. 8 (1861). (PI. 60, Fig. 4; length 34 /urn)

Barbadoes deposit, BM 2075, holotype.

This species, which belongs to Trinacria, is currently being studied by Ross & Sims (in prep.).

23. T. davyanum Grev. in Q. Jl microsc. Sci. II, 2: 232, fig. 4 (1862). (PI. 60, Fig. 5; length 157

Chimborazo Barbadoes Johnson 1862, BM 2524, holotype.

Greville states: 'The only two examples as yet known were both discovered by ... Mr. Johnson.

GREVILLE DIATOM TYPES 53

The first is a fragment [Mrs Bury] ... the other is entire [C. Johnson].' BM 2524 is the entire
valve and hence the holotype. BM 2525 is marked 'Chimborazo, Barbadoes - Johnson (Mrs
Bury) (fragment) (1862)'.
There is a detailed discussion of this species in Holmes & Brigger (1979: 172-174).

24. T. decorum Grev. in Trans, microsc. Soc. Lond. 10: 92, fig. 7 (1862). (PI. 60, Figs 7, 8;
length 58 /am)

Indian Oc. soundings Captain Pullen 2200 fathoms, BM 1922, holotype?

25. T. definitum Grev. in Trans, microsc. Soc. Lond. 12: 84, fig. 8 (1864).
Cambridge Estate Barbadoes Johnson, BM 3017, holotype.

The drawing in the Greville manuscript collection indicates BM 3017 as the holotype slide.
However, specimens have not been found.

26. T. delicatum Grev. in Trans, microsc. Soc. Lond. 9: 45 (1861); in Trans, microsc. Soc.
Lond. 9: 70, fig. 11 (1861). (PI. 60, Figs 6, 9; length 28 /am)

Barbadoes deposit, BM 2084, holotype.

27. T. denticulatum Grev. in Q. Jl microsc. Sci. II, 3: 233, fig. 14 (1863). (PI. 60, Fig. 10; length
60 /am)

Cambridge Barbadoes Johnson 1863, BM 2763, holotype.

28. T. disciforme Grev. in Q. Jl microsc. Sci. II, 3: 232, fig. 11 (1863). (PI. 60, Fig. 11; specimen
diameter 88 /am)

Cambridge Barbadoes Johnson 1862, BM 2833, holotype.

29. T. dobreeanum G. Norman ex Grev. in Trans, microsc. Soc. London. 13: 6, figs 23, 24
(1865). (PI. 61, Figs 1, 2; Valve width 42 /am, valve height from base to tips 93 /-tin, specimen
height 185 /am)

Sidney Norman 1864, BM 3340, holotype.

There is a drawing in the Greville manuscript collection but it gives no indication to which slide
was used for the published figure. The slide above is probably the specimen used for figure 24.
There is no other specimen of this taxon on the slide. That used for figure 23 has not been found.

30. T. dulce Grev. in Trans, microsc. Soc. Lond. 14: 9, fig. 20 (1866). (PI. 60, Fig. 12; length 67-5
/am)

Barbadoes Johnson 1863, BM 3094, holotype.

31. T. exornatum Grev. in Trans, microsc. Soc. Lond. 13: 9, fig. 25 (1865). (PI. 61, Figs 4-6;
length 120 /am)

Cambridge Barbadoes 1865, BM 3079, holotype.

32. T. figuratum Grev. in Trans, microsc. Soc. Lond. 13: 101, fig. 15 (1865). (PI. 61, Fig. 7;
length 25 /am)

Barbadoes Johnson 1864, BM 3281, holotype.

33. T. firmum Grev. in Trans, microsc. Soc. Lond. 12: 93, fig. 8 (1864). (PI. 61, Fig. 8; length 50
/mi)

Barbadoes Johnson 1863, BM 3087, holotype.

The slide label is not annotated by Greville. However, the original manuscript drawing reveals

that specimen MF 20/17 is the holotype.

34. T. flexuosum Grev. in Trans, microsc. Soc. Lond. 10: 93, fig. 9 (1862). (PI. 61, Fig. 3; length
58 /am)

Ind. Oc. Soundings Capt. Pullen 2200 fathoms, BM 1922 (MF 30/25).

There is no drawing of this species in the Greville manuscript collection. This is the only slide

from the type locality with a specimen of this taxon. See Simonsen (1974: 23).

35. T. foveatum Grev. in Trans, microsc. Soc. Lond. 12: 93, fig. 15 (1864). (PI. 61, Fig. 9; length
66 /am)

Cambridge Barbadoes Johnson 1863, BM 3072, holotype.

54 D. M. WILLIAMS

The holotype is situated at the extreme edge of the coverslip. The mountant is cracking and
hence good micrographs are not possible.

36. T. giganteum Grev. in Trans, microsc. Soc. Lond. 9: 77, fig. 10 (1861). (PI. 62, Fig. 1; length
150 /urn)

Barbadoes deposit (Cambridge) Mrs Bury, BM 2107, holotype.

37. T. gratiosum Grev. in Trans, microsc. Soc. Lond. 9: 85, figs 12, 13 (1864). (BM 2176, PI. 62,
Fig. 6, length 80 /am; BM 2085, PI. 62, Fig. 2, length 55 /am)

Barbadoes deposit, BM 2085; Barbadoes deposit, Norman, BM 2176, syntypes.

Two figures are given in the protologue and the drawings of both are extant in the Greville

manuscript collection. These must both be regarded as syntypes. See Holmes & Brigger (1979:

181-182).

38. T. hardmaniaiiiiiii Grev. in Trans, microsc. Soc. Lond. 13: 55, fig. 25 (1865). (PI. 62, Figs
3-5, 7; length 87 -5 /am)

Springfield Barbadoes Hardman 1864, BM 3341, holotype.

39. T. harrisonianum G. Norman & Grev. in Trans, microsc. Soc. Lond. 9: 76, fig. 9 (1861).
'Barbadoes deposit (Springfield Estate); exceedingly rare.'

There are no slides in the Greville collection from this locality that preceed 1861. It is possible
that the holotype specimen did not come from a slide in Greville's collection but from Norman's.
This slide has not been traced.

40. T. implicitum Grev. in Trans, microsc. Soc. Lond. 13: 102, fig. 25 (1865). (PI. 62, Figs 8, 9;
length 50 /urn)

Cambridge Barbadoes Johnson 1865, BM 3447, holotype.

41. T. inaequale Grev. in Trans, microsc. Soc. Lond. 12: 91, fig. 19 (1864). (PI. 63, Fig. 1;
distance between tips at widest angle 62 /am)

Cambridge Barbadoes Johnson 1863, BM 3073, holotype.

The specimen indicated on the drawing is not that which is indicated on the slide.

42. T. incoiispicuum Grev. in Trans, microsc. Soc. Lond. 9: 45 (1861); in Trans, microsc. Soc.
Lond. 9: 70, fig. 10 (1861). (PI. 63, Figs 2, 3; length 28 /am)

Barbadoes deposit, BM 2084, holotype.

This species has recently been transferred to the genus Lisitzinia (Gleser in Gleser, Dolmatova

&Lupikina, 1986:858).

43. T. inelegans Grev. in Trans, microsc. Soc. Lond. 14: 8, fig. 21 (1866). (PI. 63, Figs 4, 7;
length 71 /am)

Monterey L. H. no. 605, BM 10486, holotype.

44. T. inflatum Grev. in Q. Jl microsc. Sci. II, 3: 232, fig. 15 (1863). (PI. 63, Fig. 5; length 77-5
/am) .

Cambridge Barbadoes Johnson 1863, BM 2873, holotype.

45. T. inglorium Grev. in Trans, microsc. Soc. Lond. 13: 103, fig. 18 (1865).
'Manilla; obtained from shell-cleanings, tty L. Hardman, Esq.'

The drawing in the Greville manuscript collection is annotated with '120 H Manilla'. This
probably indicates a Hardman slide. This slide has not been traced.

46. T. inopinatum Grev. in Trans, microsc. Soc. Lond. 9: 84, fig. 10 (1861). (PI. 63, Fig. 6;
length 42-5 /am)

Barbadoes deposit, BM2081, holotype? 1 "
See Bergon (1892ft: 134).

47. T. inornatum Grev. in Trans, microsc. Soc. Lond. 10: 94, fig. 10 (1862). (PI. 63, Fig. 8;
length 55 /am)

Indian Oc. soundings 2200 fathoms, BM 2566, holotype.

GREV1LLE DIATOM TYPES 55

There is no drawing of this specimen in the Greville manuscript collection. The above slide is the
only one in Greville's collection with a specimen. See Simonsen (1979: 23).

48. T. insignis Grev. in Trans, microsc. Soc. Lond. 9: 75, fig. 5 (1861). (PI. 64, Fig. 1; length 73

Barbadoes deposit, BM 2168, holotype.

49. T. irregulare Grev. in Trans, microsc. Soc. Lond. 12: 92, fig. 5 (1864). (Micrographs not
included)

Cambridge Barbadoes Johnson 1863, BM 3074, holotype.

50. T. kittonianum Grev. in Trans, microsc. Soc. Lond. 13: 8, fig. 18 (1865). (PI. 67, Fig. 3;
length across tips at widest angle, 140 /*m)

Nottingham deposit, 4427, Wynne Baxter, BM 59049, holotype?

In the protologue, Greville refers to drawings prepared by Mr Kitton of this species which Kitton

identified as T. solenoceros Ehrenb. In Wynne Baxter's collection there are two of Kitten's

slides from Nottingham of T. solenoceros. BM 59050 is a specimen referable to T. solenoceros

but BM 59049 is a specimen of T. kittonianum. It is likely that this is the slide from which the

drawings were made and thus is the holotype. There appear to be no specimens in the Greville

collection.

51. T. labrynthaeum Grev. in Trans, microsc. Soc. Lond. 9: 45 (1861); in Trans, microsc. Soc.
Lond. 9: 70, fig. 12(1861).

Barbadoes deposit, BM 2079, holotype.

This specimen cannot be traced on the slide that is indicated as the type.

52. T. latum Grev. in Trans, microsc. Soc. Lond. 13: 103, fig. 20 (1865).
'Singapore; obtained from shell-cleanings, by L. Hardman, Esq.'

The drawing in the Greville manuscript collection is annotated with '122 H Singapore' . This slide
is probably from the Hardman collection but has not been traced.

53. T. lautum Grev. in Trans, microsc. Soc. Lond. 14: 82, fig. 20 (1866). (PI. 64, Fig. 2, 4-6;
length 135 /u,m)

Barbadoes 1865, BM 3463, holotype.

54. T. ligulatum Grev. in Trans, microsc. Soc. Lond. 12: 91, fig. 9 (1864). (PI. 64, Fig. 3;
distance between tips at widest angle, 88 /xm)

Cambridge Barbadoes Johnson 1863, BM 3088, holotype.

The slide label is not annotated with respect to T. ligulatum. However, the drawing indicates
that specimen MF 24/11 is the holotype which is marked simply 'Triceratium' 1 . The slide is made
of rather thick glass and the mountant poor, hence high magnification is not possible.

55. T. lineatum Grev. in Trans, microsc. Soc. Lond. 11: 75, fig. 24 (1863). (PI. 64, Fig. 7; length
87-5 /xm)

Cambridge Barbadoes Johnson 1863, BM 2606, holotype.

The holotype slide is of thick glass and hence high magnification is not possible.

56. T. lineolatum Grev. in Q. Jl microsc. Sci. II, 3: 233, fig. 16 (1863). (PI. 64, Fig. 8, 9; length
102-5 )am)

Bridgewater Barbadoes Johnson Jany. 1 1863, BM 2743, holotype.

The holotype slide is of thick glass and hence high magnification is not possible.

57. T. lobatum Grev. in Q. Jl microsc. Sci. II, 3: 233, fig. 13 (1863).
Cambridge Barbadoes Johnson 1863, BM 2757, holotype.

It is likely that the coverslip of this slide has moved at sometime in the past. The MF location
(28/35) given on the drawing does not locate this species, and it is not in the surrounding area.

58. T. mammosum Grev. in Trans, microsc. Soc. Lond. 14: 9, fig. 19 (1866). (PI. 64, Fig. 10;
length 47-5 /xm)

Cambridge Barbadoes Johnson 1865, BM 3434, holotype.

56 D. M. WILLIAMS

59. T. microcephalum Grev. in Trans, microsc. Soc. Lond. 9: 74, fig. 4 (1861). (PI. 64, Figs 11,
12; distance between tips at only angle, 50 /zm)

Barbadoes deposit, BM 2095, holotype.

Greville states that this species is exceedingly rare in the Barbados deposits. The holotype is

partially fragmented and obscured by debris.

60. T. microstictum Grev. in Trans, microsc. Soc. Lond. 12: 90, fig. 17 (1864). (PI. 65, Figs 1-3;
length 120 /um)

Cambridge Barbadoes Johnson 1864, BM 3033, holgrtype.

61. T. modestum Grev. in Trans, microsc. Soc. Lond. 12: 93, fig. 14 (1864). (PI. 67, Fig. 5;
length 57- 5 /um)

Barbadoes Johnson 1863, BM 3090, holotype.

62. T. moronense Grev. in Trans, microsc. Soc. Lond. 13: 32, fig. 18 (1865). (PI. 65, Figs 4, 5;
length 110 /urn)

Moron deposit Johnson 1864, BM 3236, holotype.

63. T. nebulosum Grev. in Trans, microsc. Soc. Lond. 9: 86, fig. 15 (1861).
'Barbadoes'.

The drawing in the Greville manuscript collection is annotated with 'Norman (215) - Barbados' .
This slide has not been traced.

64. T. neglectum Grev. in Trans, microsc. Soc. Lond. 13: 7, fig. 20 (1865). (PI. 65, Figs 7, 8;
length 55 /am)

Barbadoes Johnson 1863, BM 2941, holotype.

65. T. nitescens Grev. in Trans, microsc. Soc. Lond. 13: 8, fig. 19 (1865). (PI. 65, Figs 10, 11;
length 55 /xm)

Barbadoes Johnson 1864, BM 3198, holotype.

This specimen is located at the extreme edge of the coverslip and in rather thick and fragmented

mountant.

66. T. nitidum Grev. in Trans, microsc. Soc. Lond. 9: 44, fig. 13 (1861). (PI. 66, Figs 1, 2; length
30 /Am)

[Barbadoes deposit], BM 2072, holotype.

This specimen is located at the extreme edge of the coverslip and in rather thick and fragmented

mountant.

67. T. normanianum Grev. in Q. Jl microsc. Sci. II, 3: 234, fig. 19 (1863).
'Barbadoes deposit, from Cambridge estate; G. Norman, Esq.'

A drawing in the Greville manuscript collection is annotated 'no. 9 Norman'. As Greville states
that Norman was the only other person to have seen this species, the annotation probably refers
to Norman's holotype slide. This slide has not been found in BM.

68. T. notabilis Grev. in Trans, microsc. Soc. Lond. 9: 74, figs 2, 3 (1861). (PI. 67, Figs 1, 2, 4;
length 82-5 /xm)

Barbadoes deposit, BM 2086, BM 2089, BM 2094.

Although there are only two figures illustrating this species, the drawings in Greville's
manuscript collection give three slide numbers: BM 2086 (MF 28/27) and BM 2094 (MF 23/12
and 19/14) on one drawing (figure 2), and BM 2089 (MF 31/21) on the other (figure 3). These are
syntypes. BM 2089 has two additional specimens. The micrographs are taken from BM 2089
(MF 31/21).

69. T. obesum Grev. in Trans, microsc. Soc. Lond. 12: 90, fig. 11 (1864). (PI. 65, Figs 6, 9;
length 25 /u,m)

Cambridge Barbadoes Johnson 1863, BM 3078, holotype.

This slide has been broken and part of the coverslip has fragmented. Fortunately this does not

affect the specimen.

GREVILLE DIATOM TYPES 57

70. T. obscurum Grev. in Trans, microsc. Soc. Lond. 9: 76, fig. 8 (1861). (PI. 66, Fig. 3; length
51-5 /mi)

Trinidad Dr Davy, BM 2162, holotype.

This species should rightly be considered as belonging to the genus Azpeitia. The annulus and
distinctive border are clearly seen on the micrograph, although neither is apparent on Greville's
original drawing. The correct name for this species is given in the appendix.

71. T. oculatum Grev. in Trans, microsc. Soc. Lond. 12: 94, fig. 20 (1864). (PI. 66, Figs 4, 5;
length 55 /mi)

Barbadoes deposit Johnson 1863, BM 3099, holotype.

72. T. ornatum Grev. in Trans, microsc. Soc. Lond. 9. 45 (1861). (PI. 67, Fig. 6; length 47-5 /mi)
Barbadoes deposit, BM 2072, BM 2079, BM 2108, syntypes.

Greville described this species in one of the early series of the 'New and Rare . . .' papers where
he intended to publish fuller descriptions and a figure in later parts. This species was never
re-described nor was an illustration published. In the Greville manuscript collection there is a
drawing of T. ornatum. The annotations indicate BM 2079 (MF 34/14) and BM 2072 (MF 21/17).
The drawing is also annotated with an inked number '15', indicating that Greville was
considering it for publication at some stage as figure 15 of a plate.

73. T. pallidum Grev. in Trans, microsc. Soc. Lond. 12: 84, fig. 7 (1864). (PI. 66, Figs 6, 7;
length 65 /mi)

[Cambridge Estate Barbadoes Johnson], BM 2972, holotype.

There is no indication on the slide of the whereabouts of this specimen. The drawing gives the

location.

74. T. partitum Grev. in Trans, microsc. Soc. Lond. 12: 14, fig. 8 (1864). (PI. 66, Fig. 8; length
65 /xm)

Cambridge Barbadoes Johnson 1862, BM 2830, holotype.

This specimen is located on the extreme edge of the coverslip in cracked mountant, hence the

poor micrograph.

75. T. pauperculum Grev. in Trans, microsc. Soc. Lond. 13: 55, fig. 26 (1865). (PI. 66, Figs 9,
10; length 37-5 /mi)

[Cambridge Estate Barbadoes Johnson], BM 3409, holotype?

Greville's annotation on the manuscript drawing is '[BM] 3289, 24/33'. However, BM 3289 is a
slide from Arran and must have been given in error. BM 3409 has a specimen identified by
Greville as T. pauperculum, located at MF 24/33, and is probably the holotype.

76. T. pectinatum Grev. in Trans, microsc. Soc. Lond. 10: 92, fig. 6 (1862).
Scrapings of Strombus shell G. Norman, (303) 1860, BM 2484.

The above slide is the only one in Greville's collection that could be a candidate for the holotype.
No specimen has been located on the above slide. The type specimens are probably from a
G. Norman slide. These have not yet been traced.

77. T. perminutum Grev. in Trans, microsc. Soc. Lond. 12: 89, fig. 18 (1864). (PI. 68, Figs 1, 2;
length 32-5 /mi)

Barbadoes 1865, BM 3107, holotype.

78. T. perpusillum Grev. in Trans, microsc. Soc. Lond. 12: 92, fig. 13 (1864). (PI. 68, Figs 3, 6;
length 23 /mi)

Barbadoes 1865, BM 3114, holotype.

79. T. picturatum Grev. in Trans, microsc. Soc. Lond. 14: 83, fig. 19 (1866). (PI. 68, Figs 7, 8;
distance between tips at widest angle 80 /am)

Barbadoes 1864, BM 3044, holotype.

80. T. plumosum Grev. in Trans, microsc. Soc. Lond. 12: 85, fig. 4 (1864). (PI. 69, Figs 1, 4;
length 123 /xm)

58 D. M. WILLIAMS

Cambridge Estate Barbadoes, BM 2779, BM 3037, BM 3071, BM 3075, BM 3085 lectotype, BM
3096, BM 3105, BM 3108, BM 3186, BM 3441, BM 3443, BM 4778.

There is a drawing in the Greville manuscript collection but no indication as to which slide was
used for the published figure.

81. T. polygonium Grev. in Trans, microsc. Soc. Lond. 13: 105, fig. 14 (1865). (PI. 69, Figs 2, 5;
length 92-5 /xm)

Rock Ballast Stoneferry nr. Hull [4422], BM 59044, holotype?

There is a single slide in the Greville collection from the locality indicated in the protologue: BM
1898 'Stoneferry, Hull, G. Norman, 11.59 (222)'. No specimen of T. polygonium could be
located. However, in Wynne-Baxter's collection there is a slide marked 'Rock Ballast Stone-
ferry nr. Hull [4422] Triceratium polygonium Grev.' In Baxter's handwriting, and 'Figured by
Dr. Greville in J.M.S. vol. XIII, pi. VII, fig. 14.' in Kitten's own hand. As Greville says in the
protologue that his specimen came from Kitten's cabinet, it is highly probable that this is the
holotype.

82. T. praetenue Grev. in Trans, microsc. Soc. Lond. 12: 89, fig. 16 (1864). (PI. 68, Figs 4, 5;
length 42-5 /am)

Barbadoes Johnson 1863, BM 3076, holotype.

Two specimens are indicated on the slide, but the manuscript drawings indicate that the

specimen at MF 24/33 is the holotype.

83. T. productum Grev. in Trans, microsc. Soc. Lond. 9: 45 (1861); in Trans, microsc. Soc.
Lond. 9: 69, fig. 9 (1861). (PI. 69, Fig. 3; length 62-5 /-tin)

Barbadoes deposit, BM 2084, holotype.

84. T. prominens Grev. in Q. Jl microsc. Sci. II, 3: 231, fig. 10 (1863).
Cambridge Barbadoes Johnson 1863, BM 2850, holotype.

This slide is cracked and the specimen has probably moved from its original position.

85. T. pulcherrimum Grev. in Trans, microsc. Soc. Lond. 9: 82, fig. 6 (1861). (PI. 69, Figs 6, 7;
PI. 70, Fig. 1; length across longest side 102 /mi)

Barbadoes deposit (Cambridge) Mrs Bury, BM 2107, holotype.
See Holmes & Brigger (1979: 188).

86. T. quadrangulare Grev. in Trans, microsc. Soc. Lond. 13: 10, fig. 26 (1864). (PI. 70, Figs 7,
8; valve length 65 //,m)

Cambridge Estate Barbadoes Johnson 1863, BM 3186; Cambridge Estate Barbadoes Johnson

1864, BM 3190, syntypes.

The micrograph is from BM 3190 (MF 16/20).

87. T. quadratum Grev. in Trans, microsc. Soc. Lond. 13: 104, fig. 19 (1865). (PI. 70, Figs 2, 3;
length 112/xm)

Cambridge Barbadoes Johnson 1863, BM 2902, 3263 syntypes.

The manuscript drawing is annotated with two slide numbers, 2902 and 3263. Greville gives the
location MF 36/28 for the specimen on BM 2902 but this does not locate a specimen. BM 3263 is
given as a slide number only.

88. T. quadricorne Grev. in Trans, microsc. Soc. Lond. 13: 103, fig. 16 (1865). (PI. 70, Figs 4, 5;
distance across centre of valve 50 /xm)

New Caledonia 1862, BM 2767, holotype.

89. T. quinquelobatum Grev. in Trans, microsc. Soc. Lond. 14: 83, fig. 21 (1866).
'Moron deposit, Province of Seville; Rev. T. G. Stokes.'

The drawing in the Greville manuscript collection and series of manuscript notes are both
annotated 'Moron, Stokes'. There is only a single slide that corresponds to these data, but no
specimen has yet been located on the slide.

90. T. repletum Grev. in Trans, microsc. Soc. Lond. 14: 83, fig. 18 (1866). (PI. 71, Figs 1, 2;
length 70 /Am)

GREV1LLE DIATOM TYPES 59

Cambridge Barbadoes Johnson 1863, BM 3051, holotype.

91. T. reticulatum Grev. in Trans, microsc. Soc. Lond. 13: 104, fig. 21 (1865). (PI. 71, Figs 3, 4;
distance across valve centre, 65 //,m)

Barbadoes Cambridge Estate Johnson 1864, BM 3416, holotype.

The manuscript drawing identifies specimen MF 23/26 as the holotype. At that location there
appears to be a whole frustule with each valve slightly separated, giving two overlapping images.
The micrograph is of the uppermost valve.

92. T. robertsianum Grev. in Q. Jl microsc. Sci. II, 3: 231, fig. 9 (1863). (PI. 71, Figs 5, 6; length
117 /xm)

Curteis Straits, BM 2611, holotype.

93. T. robertsianum Grev. in Trans, microsc. Soc. Lond. 14: 7, fig. 22 (1866). (PI. 72, Figs 1,2;
length 85 /xm)

[Woodlark Is. South Pacific, Mr Roberts], BM 2725, holotype.

The location of the holotype specimen has been obtained from the manuscript drawing and the
slide annotated by Mr R. Ross. The slide label was not annotated by Greville.
The name is a later homonym of the above and therefore illegitimate under Art. 64.1.

94. T. robustum Grev. in Trans, microsc. Soc. Lond. 9: 71, fig. 15 (1861). (PI. 72, Figs 5, 6;
length 85 /xm)

Cove Calvert Co. Maryland, BM 2119, syntypes.

There is a drawing in the Greville manuscript collection but no indication is given of which

specimen was used for the published figure. There are three specimens on this slide.

95. T. roperianum Grev. in Trans, microsc. Soc. Lond. 10: 93, fig. 8 (1862). (PI. 72, Figs 7, 8;
length 20 /xm)

Indian Ocean soundings 2200 fathoms, BM 2534 holotype.

96. T. rotundatum Grev. in Trans, microsc. Soc. Lond. 9: 75, fig. 6 (1861). (PI. 71, Fig. 7;
distance between tips at widest angle, 40 /am)

Barbadoes deposit, BM 2111, holotype.

97. T. rylandsianum Grev. in Trans, microsc. Soc. Lond. 12: 90, fig. 6 (1864). (PI. 72, Figs 3, 4;
distance between tips at widest angle 56 /xm)

Cambridge Barbadoes Johnson 1863, BM 3074, holotype.

98. T. sexangulatum Grev. in Trans, microsc. Soc. Lond. 13: 103, fig. 24 (1865). (PI. 73, Figs 1,
2; valve length 48 /xm)

[Woodlark Island Dr Roberts], BM 2725, holotype.

99. T. smithianum Grev. in Trans, microsc. Soc. Lond. 12: 92, fig. 7 (1864). (PI. 70, Fig. 6;
length 125 /xm)

Barbadoes, 1865, BM 3105, BM 3114, BM 3469, lectotype.

Greville noted: This species was kindly communicated to me by its discoverer, Mr. G. J. Smith,
of Workington, and the specimen accidentally destroyed, but not before I had made the drawing
now engraved'. Greville says he saw specimens additional to the holotype before he published
this name. There are three specimens in his collection which are paratypes. BM 3469 is an entire
valve and designated as lectotype. The drawing is in the Greville manuscript collection.

100. T. stokesianum Grev. in Trans, microsc. Soc. Lond. 14: 8, fig. 23 (1866).
'Moron deposit, Province of Seville; Rev. T. G. Stokes.'

Greville states in the protologue: 'No one but Mr. Stokes has been so fortunate as to discover the
subject of the present notice.' The drawing is in the Greville manuscript collection but there is
only a single slide that corresponds to the locality data. No specimen has yet been located on the
slide.

101. T. subcapitatum Grev. in Q. Jl microsc. Sci. II, 3: 234, fig. 20 (1863). (PI. 73, Figs 3, 4;
distance between tips at widest angle, 48-5 /tm)

60 D. M. WILLIAMS

[Cambridge Estate Barbadoes Johnson], BM 2864, holotype.

The glass of this slide is very thick and only just allows high magnification to be used.

102. T. tesellatum Grev. in Trans, microsc. Soc. Lond. 9: 71, fig. 14 (1861). (PI. 73, Figs 5, 6;
length 50 /mi)

Nottingham deposit Maryland, [Norman], BM 2185, holotype.

There are three specimens indicated on the slide label. The manuscript drawing indicates

specimen MF 22/32 as holotype.

103. T. thwaitesianum Grev. in Trans, microsc. Soc. Lond. 10: 28, fig. 5 (1862). (PI. 73, Figs 7,
8; length 1 13 /mi)

C. B. Sp. [Cape of Good Hope] Dr Macrae 1861, BM 2393, holotype.
The holotype is one of four specimens on a selected slide.

104. T. trilineatum Grev. in Trans, microsc. Soc. Lond. 13: 56, fig. 27 (1865). (PI. 73, Figs 9, 10;
length 35 /urn)

Barbadoes Johnson 1864, BM 3272, holotype.

105. T. tumidum Grev. in Q. Jl microsc. Sci. II, 3: 234, fig. 18 (1863). (PI. 73, Figs 11, 12; length
100 /Am)

Barbadoes Johnson 1863, BM2845, holotype.

106. T. unguiculatum Grev. in Trans, microsc. Soc. Lond. 12: 85, fig. 9 (1864). (PI. 74, Figs 1-3;
length across valve face 88 /urn)

Cambridge Barbadoes Johnson 1862, BM 2806, holotype.

Information from the manuscript drawing indicates that specimen MF 33/18 is the holotype and

this has been annotated as such by Mr R. Ross.

107. T. variegatum Grev. in Trans, microsc. Soc. Lond. 9: 85, fig. 14 (1861).
'Barbadoes'.

Greville states in the protologue: 'Of this beautiful diatom I have seen only a single specimen.'
The drawing is annotated with 'Barbadoes - Norman (from Kitton).' This slide has not been
traced. See Holmes & Brigger (1979: 193).

108. T. venulosum Grev. in Trans, microsc. Soc. Lond. 12: 90, fig. 21 (1864).
Cambridge Barbadoes deposit, C. Johnson, 1863, BM 3046, holotype.

The drawing in the Greville manuscript collection is annotated with '[BM] 3046, 12/25', but no
specimen has been located.

109. T. westianum Grev. in Trans, microsc. Soc. Lond. 9: 43, fig. 11 (1861). (PI. 74, Fig. 5;
distance between tips at widest angle, 40 /mi)

Barbadoes deposit, BM 2029, holotype.

The manuscript drawing indicates BM 2028 as the holotype but no specimen is located.
However, the MF co-ordinates provided reveal a specimen on BM 2029 and 'hence this is
considered to be the holotype and Greville's annotation a typographical error.

110. T. zonatulatum Grev. in Trans, microsc. Soc. Lond. 13: 102, fig. 17 (1865). (PI. 74, Figs 7,
8; distance across valve centre 30 /mi; PI. 74, Fig. 6, distance across valve face 30 /mi)
Singapore, BM 10523, L. H. no. 645, holotype.

As indicated in Greville's protologue, the slide used for the type description came from L.
Hardman. The drawing is marked '122H Singapore'. The number on Greville's drawing, 122,
bears no relationship to Hardman's numbering. On the slide there are 15 specimens. All must be
regarded as syntypes.

111. T. zonatum Grev. in Trans, microsc. Soc. Lond. 12: 84, fig. 3 (1864). (PI. 74, Fig. 4; length
60 /am)

Cambridge Barbadoes Johnson 1864, BM 3040, holotype.

The slide is made from thick glass and hence high magnification cannot be used.

GREVILLE DIATOM TYPES 61

LXXIX. XANTHIOPYXIS Ehrenb.

1. X? umbonatus Grev. in Trans, microsc. Soc. Lond. 14; 2, fig. 5 (1866). (PI. 74, Figs 9-11;

specimen diameter 95 ^im)

Monterey, BM 10571, L. H. no. 672, BM 10573, L. H. no. 673.

There is a drawing in the Greville manuscript collection but it is without annotations. The above

specimens are considered syntypes.

Appendix

The nomenclatural changes that have been made in this paper are detailed below.

1. Azpeitia obscurum (Grev.) P. A. Sims, comb, nov.*

Triceratium obscurum Grev. in Trans, microsc. Soc. Lond. 9: 76 (1861).

Triceratium antiquum Pant, in Beitr. foss. Bacill. Ungarns 1: 51 (1886).

Azpeitia temperei M. Perag. in Temp. & Perag., Dial. Monde entier, 326, fasc. 21, nos 658-660

(1912).

Azpeitia antiqua (Pant.) P. A. Sims in Syst. Bot. Monogr. 13: 6 (1986).
For discussion see p. 57.

2. Odontella hastata (Grev.) J. Fenner ex D. M. Williams, comb. nov.
Hemiaulus hastatus Grev. in Trans, microsc. Soc. Lond. 13: 31, fig. 15 (1865).

Fenner (1982) transferred H. hastatus to Odontella, but the combination is not effectively
published as it appeared in an unpublished Ph. D. thesis. See p. 36.

3. Perissonoe parvula (Grev.) D. M. Williams, comb. nov.

Amphitetras parvula Grev. in Edinb. New phil. J. II, 18: 37, fig. 12 (July 1863); in Trans, hot.

Soc. Edinb. 7: 537, fig. 12 (1863) [non A parvula Janisch & Rabenh. in Beitr. kenntn. Alg. : 4,

pi. 1, fig. 4, (Oct.-Nov. 1863).]

Amphitetras cruciata Janisch & Rabenh. in Beitr. kenntn. Alg. : 4, pi. 1, fig. 5 (Oct.-Nov. 1863).
Amphitetras crucifera Kitton in Sci. Gossip 3: 271, fig. 205 (1867).
Rhaphoneis amphiceros var. tetragona Grun. in Van Heurck, Syn. Dial. Beige: pi. 116, fig. 16

(1883).

Amphitetras grevillei De Toni, in Syll. Alg. 2: 906 (1894), nom. superfl. (Art. 63.1).
Rhaphoneis amphiceros var. cruciata (Janisch & Rabenh.) Mereschk. in Scripta Bot. Horti Univ.

Imp. Petropol. 18: 151, pi. 4, figs 19, 20 (1902).
Perissonoe cruciata (Janisch & Rabenh.) G. W. Andrews & Stoelzel in Proc. 7th Internal. Dial.

Symp. : 226, pi. 1, figs 1-8, pi. 2, figs 21, 23-26, pi. 3, figs 27, 28, 30, pi. 4, figs 31-34 (1984).

When Janisch & Rabenhorst (1863) described the new species Amphitetras cruciata they were
unaware of Greville's slightly earlier publication of the same species with the name A. parvula.
Simultaneously, Janisch & Rabenhorst published another new Amphitetras under the name of
A. parvula. Their A. parvula is clearly different from that of Greville's. The exact date of
publication of the Janisch & Rabenhorst taxon is Oct.-Nov. 1863 (according to Stafleu &
Cowan, 1983). Greville published his name in both the Transactions of the Botanical Society of
Edinburgh and The Edinburgh New Philosophical Journal. The latter was published in July 1863
and hence predates that of Janisch & Rabenhorst. De Toni (1894) proposed a new name, A.
grevillei, for A. parvula Grev., but this is a nom. superfl. Greville also made reference to A.
crucifera Kitton. This name, however, remained unpublished until 1867. A slide of Kitton's
material is present in the Greville herbarium (BM 4412). See p. 7.

* This note is by P. A. Sims.

62 D. M. WILLIAMS

4. Pseudauliscus grevillei R. Ross, nom. nov.*

Auliscus nebulosus Grev. in Trans, microsc. Soc. Land. 11: 74, fig. 21 (1863).

Pseudauliscus nebulosus (Grev.) Rattray inJIR. microsc. Soc. 1888: 904 (1888), non Pseudaulis-
cus nebulosus Leuduger-Fortmorel in Mem. Soc. Emul. Cotes Nord: 64, pi. 7, fig. 74 (1879)
(Art. 64.1). Seep. 17.

Acknowledgements

I am extremely grateful to Pat Sims for encouraging me to continue and complete this project and for her
detailed comments on the manuscript. I am indebted to R. Ross for his advice on nomenclatural matters
and for his comments on the manuscript. Barrie Paddock and Alan Eddy also provided useful remarks. M.
Sullivan provided details of his published and unpublished studies on Greville types. I thank David Sutton
for assistance in planning this project. Finally, I am grateful to the staff of the Photographic Unit for
processing and printing the large number of micrographs.

References and bibliography

Andrews, G. W. 1986. Morphology and stratigraphic occurence of the marine diatom genus Brightwellia

Ralfs. In M. Ricard (Ed.), Proceedings of the Eight International Diatom Symposium: 125-140,

Koenigstein.
& Stoelzel, V. A. 1984. Morphology and evolutionary significance oiPerissonoe, a new marine diatom

genus. In D. G. Mann (Ed.), Proceedings of the Seventh International Diatom Symposium: 225-240,

Koenigstein.

Barker, J. W. & Meakin, S. H. 1944. New and rare diatoms. /. Quekett microsc. Club IV, 2: 18-22.
Bergon, P. 1892a. Monographic du genre Entogonia. Diatomiste 1: 83-90.

- 18926. Monographic du genre Entogonia. (fin). II. Classification et description des especes.

Diatomiste 1: 128-147.
Bone, E. C. F. 1969. Glyphodiscus stellatus Greville. Note on the dissimilar nature of the component valves

of the frustules. /. Quekett microsc. Club 31: 113-114.
Burke, J. F. 1970-1974. A review of the genus Aulacodiscus. The Staten Island Institute of Arts and

Sciences, Staten Island, New York. Fascicules 7 & 8.
& Woodward, J. B. 1963-1970. A review of the genus Aulacodiscus. The Staten Island Institute of

Arts and Sciences, Staten Island, New York. Fascicules 1-6.
Cox, E. J. 1975a. A reappraisal of the diatom genus Amphipleura Kiitz. using light and electron

microscopy. Br. phycol. J. 10: 1-12.

1975ft. Further studies on the genus Berkeleya Grev. Br. phycol. J. 10: 205-217.

1982. Taxonomic studies on the diatom genus Navicula Bory. IV. Climaconeis Grun., a genus

including Okedenia inflexa (Breb.) Eulenst. ex De Toni and members of Navicula sect, johnsonieae

sensu Hustedt. Br. phycol. J. 17: 147-168.

1983. Observations on the diatom genus Donkinia Ralfs in Pritch. III. Taxonomy. Botanica mar. 26:

567-580.

De Toni, G. B. 1894. Sylloge algarum 2 (3): 819-1556. Patavii.
Ehrenberg, C. G. 1856. Mikrogeologie. Das Erde und felsen schaffende Wirken des unsichtbar kleinen

selbststdndigen Lebens aufder Erde. Fortsetzung.
Fenner, J. 1982. Diatoms in the Eocene and Oligocene sediments off N. W. Africa, their stratigraphic and

paleographic occurences. Ph. D. dissertation, University of Kiel, West Germany.
Fryxell, G. A., Sims, P. A. & Watkins, T. P. 1986. Azpeitia (Bacillariophyta): Related genera and

promorphology. Syst. Bot. Monogr. 13.
Gleser, Z. I. 1975. K revizii roda Triceratium Ehr. sensu Hustedt, 1930 (Bacillariophyta). [Towards a

revision of the genus Triceratium Ehr. sensu Hustedt (Bacillariophyta).] Bot. Zh. SSSR 60: 1304-1310.
1986. O polozhenii roda Triceratium s.str. (Bacillariophyta) v sisteme diatomovykh vodoroslei. [On

the position of the genus Triceratium s.str. (Bacillariophyta) in the diatom system.] Bot. Zh. SSSR 71:

1543-1545.

Dolmatova, L. M. & Lupikina, E. G. 1986. Marine paleogenic diatoms algae from eastern

Kamchatka. Bot. Zh. SSSR 71: 851-859.
Gombos, A. M. 1980. The early history of the diatom family Asterolampraceae. Bacillaria 3: 227-272.
1982. Early and middle Eocene diatom evolutionary events. Bacillaria 5: 225-242.

* This note is by R. Ross.

GREVILLE DIATOM TYPES 63

1983. Middle Eocene diatoms from the south Atlantic. Initial Rep. deep Sea Drilling Proi. 71:

565-581.
& Ciesielski, P. F. 1983. Late Eocene to early Miocene diatoms from the southwest Atlantic. Initial

Rep. deep Sea Drilling Pro]. 71: 583-634.
Greville, R. K. 1822. A description of two new plants of the order Algae, found in Scotland. Mem.

Wernerian nat. Hist. Soc. 4: 213-217.
1827. Scottish cryptogamic flora, or coloured figures and descriptions of cry ptogamic plants, belonging

chiefly to order Fungi. 1822-1828. Edinburgh.
- 1833. The English flora of Sir J. E. Smith by Williams J. Hooker, 1824-1836; vol. 5, Div. IV.

Diatomaceae: 401-415.
1855. Report on a collection of Diatomacae made in the district of Braemar by Professor Balfour and

Mr. George Lawson. Ann. Mag. nat. Hist. II, 15: 252-261. [Also published in Trans, hot. Soc. Edinb. 5:

45-54.]

1857a. Notice of a new genus of Diatomaceae. In W. Gregory, On new forms of marine Diatomaceae

found in the Firth of Clyde and in Loch Fine. Trans. R. Soc. Edinb. 21: 536-538.

1857ft. Descriptions of some new diatomaceous forms from the West Indies. O. Jl microsc. Sci. 5:
7-12.

1859a. Descriptions of new species of British Diatomaceae chiefly observed by the late Professor
Gregory. Q. Jl microsc. Sci. 7: 79-86.

1859ft. Descriptions of Diatomaceae observed in Californian Guano. Q. Jl microsc. Sci. 7: 155-166.

1859c. Descriptions of some new species and varieties of Naviculae, etc., observed in Californian
Guano. Edinb. Newphil. J. 10: 25-30; Trans, hot. Soc. Edinb. 6: 245-249.

lS59d. On Plagiogramma, a new genus of Diatomaceae. Q. Jl microsc. Sci. 7: 207-211.

1860o. On Campy iodiscus. Trans, microsc. Soc. Land. 8: 29-32.

1860ft. A monograph of the genus Asterolampra, including As teromphalus and Spatangidium. Trans,
microsc. Soc. Lond. 8: 102-124.

1861a. Descriptions of new and rare diatoms. Series I. Trans, microsc. Soc. Lond. 9: 39-45.

1861ft. Descriptions of new and rare diatoms. Series II. Trans, microsc. Soc. Lond. 9: 67-73.

1861c. Descriptions of new and rare diatoms. Series III. Trans, microsc. Soc. Lond. 9: 73-77.

lS6ld. Descriptions of new and rare diatoms. Series IV. Trans, microsc. Soc. Lond. 9: 79-87.

1862a. Descriptions of new and rare diatoms. Series V. Trans, microsc. Soc. Lond. 10: 18-29.

1862ft. On the Asterolamprae of the Barbadoes deposit. Trans, microsc. Soc. Lond. 10: 41-55.

1862c. Descriptions of new and rare diatoms. Series VI. Trans, microsc. Soc. Lond. 10: 89-96.

1862d. Descriptions of new and rare diatoms. Series VII. Q. Jl microsc. Sci. II, 2: 231-236.

1863a. Descriptions of new and rare diatoms. Series VIII. Trans, microsc. Soc. Lond. 11: 13-21.

1863ft. A monograph of the genus Auliscus. Trans, microsc. Soc. Lond. 11: 36-53.

1863c. Descriptions of new and rare diatoms. Series IX. Trans, microsc. Soc. Lond. 11: 63-76.

1863d. Descriptions of new and rare diatoms. Series X. Q. Jl microsc. Sci. II, 3: 227-237.

1863e. Descriptions of new genera and species of diatoms from the South Pacific. Edinb. Newphil. J.
II, 18: 34-43. [Also published in Trans, hot. Soc. Edinb. 7: 534-543.]

1863/. Descriptions of new genera and species of diatoms from the South Pacific. Part 2. Edinb. New

phil. J. II, 18: 181-187. [Also published in Trans, hot. Soc. Edinb. 7: 574-580.]

1864a. Descriptions of new and rare diatoms. Series XI. Trans, microsc. Soc. Lond. 12: 8-14.

1864ft. Descriptions of new and rare diatoms. Series XII. Trans, microsc. Soc. Lond. 12: 81-86.

1864c. Descriptions of new and rare diatoms. Series XIII. Trans, microsc. Soc. Lond. 12: 87-94.

1865a. Descriptions of new and rare diatoms. Series XIV. Trans, microsc. Soc. Lond. 13: 1-10.

1865ft. Descriptions of new and rare diatoms. Series XV. Trans, microsc. Soc. Lond. 13: 24-34.

1865c. Descriptions of new and rare diatoms. Series XVI. Trans, microsc. Soc. Lond. 13: 43-57.

I865d. Descriptions of new and rare diatoms. Series XVII. Trans, microsc. Soc. Lond. 13: 97-105.

I865e. Descriptions of new genera and species of diatoms from Hong Kong. Ann. Mag. nat. Hist. Ill,
16: 1-7.

1866a. Descriptions of new genera and species of diatoms from the South Pacific. Part III. Trans, hot.
Soc. Edinb. 8: 233-238.

1866ft. Descriptions of new and rare diatoms. Series XVIII. Trans, microsc. Soc. Lond. 14: 1-9.

1866c. Descriptions of new and rare diatoms. Series XX [XIX]. Trans, microsc. Soc. Lond. 14:
77-86.

1866d. Descriptions of new and rare diatoms. Series XX. Trans, microsc. Soc. Lond. 14: 121-130.

1866e. Descriptions of new and rare diatoms from the tropics and Southern Hemisphere. Trans, hot.
Soc. Edinb. 8: 436-441.

64 D. M. WILLIAMS

Hanna, G. D. 1931. Diatoms and silicoflagellates of the Kreyenhagen Shale. Rep. St. Miner. Calif.:

187-201.

& Brigger, A. L. 1970. Observations on Liostephania. Beih. nov. Hedwigia 31: 89-100.
Hartley, B. 1986. A check-list of the freshwater, brackish and marine diatoms of the British Isles and

adjoining coastal waters./, mar. biol. Ass. U.K. 66: 531-610.
Hasle, G. R. 1973. Morphology and taxonomy of Skeletonema costatum (Bacillariophyceae). Norw. J. Bot.

20: 109-137.
- & Sims, P. A. 1985. The morphology of the diatom resting spores Syringidium bicorne and

Syringidium simplex. Br. phycol. J. 20: 219-225.

1986. The diatom genus Coscinodiscus Ehrenb.: C. argus Ehrenb. and C. radiatus Ehrenb.

Botanicamar. 29: 305-318.

& Syversten, E. E. 1980. The diatom genus Cerataulina: morphology and taxonomy. Bacillaria 3:

79-113.
Hendey, N. 1. 1969. Pygupyxis, a new genus of diatoms from a South Atlantic eocene core. Occ. Pap. Calif.

Acad. Sci. 72.
& Sims, P. A. 1987. Examination of some fossil Eupodiscoid diatoms with descriptions of two new

species of Craspedoporus Greville. Diatom Res. 2: 23-34.
Holmes, R. W. & Brigger, A. L. 1977. The morphology and occurence of Entogonia from an eocene

locality in the Indian Ocean. Beih. nov. Hedwigia 54: 215-241.

1979. The marine fossil diatom genus Entogonia Greville - A review. Bacillaria 2:

155-214.
Hustedt, F. 1957. Die Diatomeenflora des Flusssystems der Weser im Gebiet der Hansestadt Bremen.

Abh. naturw. Ver. Bremen 34: 181 r 440.
Janisch, C. & Rabenhorst, L. 1863. Uber Meeres-Diatomaceen von Honduras, vonJanisch undDrLudwig

Rabenhorst, 'Beitrage zurndheren kenntnis und Verbreitung derAlgen'. 1.
Jouse, A. P. 1978. Novi iskopaemiii rod diatomei Lisitzinia (Bacillariophyta). Morskaya Mikropaleont.:

47-48.
Karsten, G. 1905. Das Phytoplankton des Antarktischen Meeres nach dem Material der Deutschen

Tief see-Expedition 1898-1899. Wissenschaftliche Ergebnisse der Deutschen Tief see- Expedition aufdem

Dampfer 'Valdivia' 1898-1899. Jena.

Kitton, F. 1867. The genus Amphitetras. Hardwicke's Sci. Gossip 3: 271-272.
Krotov, A. I. & Shibkova, K. G. 1959. Species novae Diatomacearum e Paleogeno Montium Uralensium.

Bot. mat. 12: 112-129.

Lauder, W. L. 1865. On the diatomaceae of Otago, New Zealand. Bot. J. Linn. Soc. 9: 129-134.
LeCohu, R. 1983. Observations sur deux especes de diatomees du genre Diatomella: Diatomella hustedtii

Manguin et Diatomella oenkoana Maillard. Cryptogamie Algol. 4: 63-71.
Leuduger-Fortmorel, G. 1879. Catalogue des Diatomees de File Ceylan. Mem. Soc. Emul. Cotes Nord,

73 pp., 9 pis.
McVaugh, R., Ross, R. & Stafleu, F. A. 1968. An annotated glossary of botanical nomenclature. Regnum

Vegetabile 56: 1-31.
Nikolaev, V. A. 1984. De Hemidiscaceis (Hendey) Simonsen in Antarctica inventis. Nov. Sist. Nizsh. Rast.

21: 35-38.
Paddock, T. B. & Sims, P. A. 1980. Observations on the marine diatom genus Auricula and two new

genera, Undatella and Proboscidea. Bacillaria 3: 161-196.
Payne, F. W. 1922. Liostephania and its allies. London.
Pritchard, A. 1861. A history of Infusoria. 4th Ed. London.
Rattray, J. 1888o. A revision of the genus Auliscus Ehrb. and of some allied genera. // R. microsc. Soc.

1888: 861-920.

1888ft. A revision of the genus Aulacodiscus Ehrb. // R. microsc. Soc. 1888: 337-382.

1890. A revision of the genus Coscinodiscus, and some allied genera. Proc. R. Soc. Edinb. 16:

449-692.

Ross, R. 1972. On some fossil species ofHemiaulus. Trans, hot. Soc. Edinb. 41: 323-342.
1976. Some Eocene diatoms from South Atlantic cores. Part. II. Rutilaria Grev. Occ. Pap. Calif.

Acad. Sci. 123: 21-27.
& Sims, P. A. 1970. Studies of Aulacodiscus with the scanning electron microscope. Beih. nov.

Hedwigia 31: 49-88.

1971. Generic limits in the Biddulphiaceae as indicated by the scanning electron microscope.

In V. H. Heywood (Ed.), Scanning electron microscopy, systematic and evolutionary applications:
155-177, London.

GREVILLE DIATOM TYPES 65

1980. Syringidium Ehrenb., Dextrodonator Ross and Sims, nov. gen. and Abas Ross and- Sims,

nov. gen. Bacillaria 3\ 115-127.

1985. Some genera of the Biddulphiaceae (diatoms) with interlocking spines. Bull. Br. Mus. nat.

Hist. (Bot.) 13: 277-381.
Schoeman, F. R. & Archibald, R. E. M. 1986a. Observations on Amphora species (Bacillariophyceae) in

the British Museum (Natural History). I. Some species from the subgenus Oxy amphora Cleve. Nova

Hedwigia 43: 113-127.
1986ft. Observations on Amphora species (Bacillariophyceae) in the British Museum (Natural

History). II. Some species from the subgenus Psammamphora Cleve. Nova Hedwigia 43: 473-484.

1986c. Observations on Amphora species (Bacillariophyceae) in the British Museum (Natural

History). V. Some species from the subgenus Amphora. S. Afr. J. Bot. 52: 425-437.

1987a. Observations on Amphora species (Bacillariophyceae) in the British Museum (Natural

History). III. Two species from the subgenus Amblyamphora Cleve. Nova Hedwigia 44: 125-135.

1987ft. Observations on Amphora species (Bacillariophyceae) in the British Museum (Natural

History). VI. Some species from the subgenus Halamphora Cleve. Nova Hedwigia 44: 377-398.

& Sims, P. A. 1986. Observations on Amphora species (Bacillariophyceae) in the British

Museum (Natural History). IV. Some species from the subgenus Diplamphora Cleve. Cryptogamie

Algol. 7: 9-21.

Schmidt, A. 1881. Atlas der Diatomaceenkunde 17, 18. Leipzig.
Schrader, H. J. & Fenner, J. 1976. Norwegian Sea Cenozoic biostratigraphy and taxonomy. Part 1.

Norwegian Sea Cenozoic diatom biostratigraphy. Inlt. Rep. deep Sea Drilling Proj. 38: 921-1099.
Simonsen, R. 1968. Descriptions of new and rare diatoms by R. K. Greville. In R. Simonsen (Ed.), Biblthca

phycol. 6.
1974. The diatom plankton of the Indian Ocean Expedition of RV 'Meteor' 1964-1965. Meteor

ForschErgebn. , D, 19: 1-66.
1975. On the pseudonodulus of the centric diatoms, or Hemidiscaceae reconsidered. Bieh. nov.

Hedwigia 53: 83-94.
1979. The diatom system: ideas on phylogeny. Bacillaria 2: 9-71.

1982. Note on the diatom genus Charcotia M. Peragallo. Bacillaria 5: 101-116.

Stafleu, F. A. & Cowan, R. S. 1983. Taxonomic literature 2nd ed. Den Haag.

Stidolph, S. R. 1985. Occurrence of the diatom Glyphodiscus stellatus Greville living in New Zealand

coastal waters. Nova Hedwigia 41: 495-504.
Stosch, H. A. V. 1986. Some marine diatoms from the Australian Region, especially from Port Phillip Bay

and tropical north-eastern Australia. II. Survey of the genus Palmeria and the family Lithodesmiaceae

including the new genus Lithodesmioides. Brunonia 9: 29-87.
Strel'nikova, N. I. 1974. Diatomei Pozdnego Mela (Zapadnaya Sibir') [Diatoms of the Late Cretaceous

(western Siberia)]. Moskva.
Sullivan, M. J. 1986. A light and scanning microscope study of Eupodiscus radiatus Bailey (Eupodis-

caceae). In M. Ricard, Proceedings of the 8th International Diatom Symposium: 113-123. Paris.
1987 . A light and scanning electron microscope study of the marine diatom Auliscus pruinosus Bailey

(Eupodiscaceae). Br. phycol. J. 22: 33-42.

in press. A morphological study of the marine diatom Glyphodesmis eximia, the type of its genus.

Dial. Res.

Sundstrom, B. G. 1986. The marine diatom genus Rhizosolenia. A new approach to the taxonomy. Lund.
Swatman, C. C. 1948. Note on the genus Fenestrella Greville. JIR. microsc. Soc. 68: 51-54.
Van Heurck, H. 1896. A treatise on the Diatomaceae. Translated by W. E. Baxter. London.
VanLandingham, S. L. 1968. Catalogue of the fossil and Recent genera and species of diatoms and their

synonyms. Part II. Bacteriastrum through Cosdnodiscus: 494-1086. Lehre.
Wahrer, R. J., Fryxell, G. A. & Cox, E. J. 1985. Studies in pennate diatoms: Valve morphologies of

Licmophora and Campy lostylus. J. Phycol. 21: 206-217.
Whiting, M. C. & Schrader, H. 1985. Actinocyclus ingens Rattray: reinvestigation of a polymorphic

species. Micropaleontology 31: 68-75.
Williams, D. M. 1985. Morphology, taxonomy and inter-relationships of the ribbed araphid diatoms from

the genera Diatoma and Meridian (Diatomaceae: Bacillariophyta). Biblioth. diatomol. 8.
Wise, F. C. 1952. Notes on four oligocene diatoms from Oamaru, New Zealand. Ann. Mag. nat. Hist. XII,

5: 404-408.

Index

Although the catalogue is presented in alphabetical order an index has been provided to take account of
currently accepted names. Many of the taxa Greville described have undergone taxonomic changes and, in
some cases, there is still no agreement on their systematic position. This index is therefore intended only as
a guide to Greville's types and does not necessarily reflect current nomenclature.
Accepted names are in roman and synonyms in italic; new names are in bold, as are principal references.

Achnanthes angustata Grev. (= A. brevipes var.

angustata (Grev.) Cleve) 3, PI. 1, Fig. 1
carmichaelii Grev. (=A. longipes Agardh) 3, PI. 1,

Figs 2-4
costata Grev. (non A. costata Johnston 1860) 3, PI.

1, Figs 8- 10

gregoriana Grev. 3, PI. 1, Figs 11-12
pennaeformis Grev. (=A. brevipes var.

pennaeformis (Grev.) Cleve) 3, 4, PI. 1, Figs 5-7
Actinodiscus barbadensis Grev. 4, PI. 2, Fig. 1
Actinoptychus minutus Grev. 4
Amphiprora brebissoniana Grev. 4, PI. 2, Figs 2, 4, 5
chinensis Grev. 4, PI. 2, Fig. 6
?clepsydra Grev. 4, PI. 2, Fig. 8
conspicua Grev. 5, PI. 3, Figs 1, 2
delicatula Grev. (=Plagiotropis lepidoptera var.

delicatula (Grev.) Nizamuddin) 5, PI. 3, Figs 3-5
eximia Grev. 5, PI. 3, Fig. 7
hyalina Grev. (=Thalassiophysa hyalina (Grev.)

Paddock & P. A. Sims) 5, 8, PI. 2, Fig. 3
jolisiana Grev. 5, PI. 2, Fig. 7
kuetzingiana Grev. 5, PI. 4, Figs 1, 2
lata Grev. 5, PI. 4, Figs 3, 4
lepida Grev. 5, PI. 3, Figs 8, 9
lineata Grev. (=Undatella lineata (Grev.) Paddock

& P. A. Sims) 5, 8, PI. 4, Figs 8, 9
meneghiniana Grev. 6, PI. 4, Figs 7, 8
nitida Grev. 6, PI. 5, Figs 1,2
oblonga Grev. 6, PI. 3, Figs 3-5
Iparadoxa Grev. (=Tropidoneis concerta (Lewis)

Cleve) 6, PI. 5, Fig. 7
?rabenhorstiana Grev. 6, PI. 5, Fig. 8
rectangularis Grev. 6, PI. 5, Fig. 6
?superba Grev. 6, PI. 6, Figs 1-3
thwaitesiana Grev. 6, PI. 5, Figs 9, 10
?undulata Grev. 6
venusta Grev. 6

Amphitetras crudata Janisch & Rabenh. 61
crucifera Kitton 61
elegans Grev. (=Triceratium elegans (Grev.) Grun.)

7, PI. 6, Figs 4-8
grevillei De Toni 61
minuta Grev. (=Triceratium antediluvium var.

minutum (Grev.) F. W. Mills) 7, PI. 6, Fig. 5
nobilis Grev. (=Triceratium nobilis (Grev.) F. W.

Mills) 7

parvula Grev. 1, 7, 61, PI. 6, Figs 6, 7
producta Grev. 7, PI. 6, Fig. 9
punctata Grev. ( = ?Triceratium punctata (Grev.) F.

W. Mills) 7, PI. 6, Fig. 10
radiata Grev. (= Triceratium radiatum (Grev.) M.

Perag.)7,Pl. 7, Figs 1,2
Amphora flexuosa Grev. (= A. sarniensis var. flexuosa

(Grev.) Cleve) 7, PI. 7, Fig. 5
magnified Grev. (=Undatella magnifica (Grev.)

Paddock & P. A. Sims) 7, PI. 7, Figs 8, 9

pulchra Grev. ( = Undatella lineata (Grev.) Paddock

&P. A. Sims) 8, PI. 7, Figs 6, 7
sarniensis Grev. (=A. sarniensis var. flexuosa

(Grev.) Cleve) 8
sinuata Grev. 8, PI. 7, Fig. 4
undulata Grev. 8, PI. 7, Fig. 3
Arachnoidiscus grevilleanus Hardman ex Grev. 8
Asterionella synedraeformis Grev. (=Thalassiothrix

frauenfeldii (Grun.) Grun.) 8
Asterolampra aemulans Grev. 9, PI. 8, Figs 1, 2
affinis Grev. 9, PI. 8, Figs 3-6
?alienaGrev. 9, P1.9, Fig. 1
ambigua var. a Grev. 9
ambigua var. b Grev. 9, PI. 9, Fig. 2
brebissoniana Grev. 9
concinna Grev. 9, PI. 9, Figs 5, 7
crenata Grev. 10, PI. 9, Fig. 6
dallasiana Grev. 10
decora Grev. (= Asteromphalus decorus (Grev.)

Bukry & Foster) 10, PI. 10, Figs 1-4
decorata Grev. 10, PI. 10, Figs 5-8
dubia Grev. 9, 10
eximia Grev. 10, PI. 10, Figs 9, 10
hiltoniana Grev. (= Asteromphalus hiltonianus

(Grev.) Ralfs) 10, PI. 11, Figs 1, 2
kittoniana Grev. 9, 11, PI. 11, Fig. 3
laevisGrev. 9, 11, PI. ll,Figs4,5
moronensis Grev. (= Asteromphalus moronensis

(Grev.)Rattray)ll
pulchra Grev. 9, 11, PI. 11, Figs 6, 7
punctata Grev. 11, PI. 11, Figs 8, 9
ralfsiana Grev. 11, PI. 12, Fig. 1
roperiana Grev. (= Asteromphalus roperianus

(Grev.) Ralfs) 11, PI. 12, Fig. 2
rotula Grev. (= Asterolampra grevillei (Wall.)

Grev.) 12

rylandsiana Grev. 12, PI. 12, Fig. 3
scutula Grev. 9, 12, PI. 12, Fig. 4
shadboltiana Grev. (= Asteromphalus shadboltianus

(Grev.) Ralfs) 12, PI. 12, Fig. 5
simulans Grev. 9, 12, PI. 12, Fig. 6
splendida Grev. (=A. vulgaris var. planior Rattray)

13
stellata Grev. (- Asteromphalus stellatus (Grev.)

Ralfs) 12

stellulata Grev. 9, 12, PI. 12, Fig. 7
variabilis Grev. (= Asterolampra grevillei (Wall.)

Grev.) 12, PI. 9, Figs 3-4; PI. 12, Figs 8-12; PI. 13,

Figs 1,2
vulgaris var. a Grev. (= A. vulgaris Grev.) 13, PI. 13,

Figs 3-9
vulgaris var. b Grev. (= A. vulgaris Grev.) 13, PI. 13,

Fig. 10

vulgaris var. c Grev. 13
vulgaris var. d Grev. (= A. vulgaris var. cellulosa

Rattray), 13, PI. 13, Figs 11, 12

GREVILLE DIATOM TYPES

67

Asterolampra - cont.
vulgaris var. e Grev. (= A. vulgaris var. planior

Rattray) 13, PI. 14, Fig. 6
wallichiana Grev. (=Asteromphalus wallichianus

(Grev.) Ralfs) 13, PI. 14, Fig. 5
Asteromphalus elegans Grev. (= Asterolampra elegans

(Grev.) Grev.) 13, PI. 14, Fig. 4
Aulacodiscus amoenus Grev. 13, PI. 14, Fig. 3
angulatus Grev. 14, PI. 14, Fig. 1
attenuatus Rattray 14, PI. 15, Figs 10, 1 1
cinctus Grev. ex Rattray 14, PI. 15, Figs 6, 9
decorus Grev. 14, PI. 15, Figs 7, 8
elegans Grev. nom. herb. 14
extans Grev. (=A. mammosus var. extans (Grev.)

Rattray) 14, PI. 15, Figs 1,2
gigas Grev. 15, PI. 15, Fig. 4
grevilleanus G. Norman 13, 15
inflatus Grev. 15, PI. 16, Figs 1, 2, 4
jonesianus Grev. (= A. scaber var. jonesiana (Grev.)

Rattray) 15, PI. 15, Fig. 3
kilkellyanus Grev. 15, PI. 16, Fig. 5
macraeanus Grev. 15, PI. 16, Fig. 3
mammosus Grev. 15, PI. 16, Figs 7, 8
orientalis Grev. 15, PI. 17, Figs 1-3
pallidusGrev. 15
Iparadoxus Grev. (=Actinoptychus undulatus (J. W.

Bail.) Ralfs) 15, PI. 16, Fig. 6
pellucidus Grev. 15, PI. 17, Figs 4, 5
radiatus Grev. 16
sparsus Grev. (= A. kilkellyanus var. sparsa (Grev.)

Rattray) 16, PI. 17, Fig. 6
spectabilis Grev. 16, PI. 18, Figs 1,2
umbonatus Grev. 16, PI. 17, Fig. 7
Auliscus ambiguus Grev. 16, PI. 18, Figs 3, 4
australiensis Grev. 16
barbadensis Grev. 16, PI. 18, Fig. 5
cellulatus Grev. (=Pseudauliscus ralfsianus (Grev.)

Rattray) 16, PI. 18, Figs 7, 8
elaboratus Ralfs ex Grev. 16, PI. 18, Fig. 6
elegans Grev. 17, PI. 19, Figs 1,4
hardmanianus Grev. 17, PI. 19, Fig. 3
johnsonianus Grev. 17, PI. 19, Fig. 5
macraeanus Grev. 17, PI. 19, Fig. 7; PI. 20, Fig. 1
mirabilis Grev. 17
moronensis Grev. 17, PI. 19, Fig. 6
nebulosus Grev. 1, 17, 62, PI. 19, Fig. 8; PI. 20, Fig. 2
normanianus Grev. 17, PI. 20, Fig. 3
notatus Grev. 17, PI. 20, Fig. 4
ornatus Grev. 18, PI. 20, Figs 6, 7
parvulus Grev. 18, PI. 20, Fig. 5
racemosus Ralfs ex Grev. 18, PI. 21, Figs 1, 4, 5
ralfsianus Grev. ( = Pseudauliscus ralfsianus (Grev.)

Rattray) 18, PI. 21, Fig. 6
reticulatus Grev. 18, PI. 21, Figs 2, 3
Azpeitia antigua (Pant.) P. A. Sims 61
obscurum (Grev.) P. A. Sims 1, 61
temper ei M. Perag. 61
Berkeleya fragilis Grev. 18, PI. 21, Fig. 7
Biddulphia barbadensis Grev. 18, PI. 22, Figs 7-9
chinensis Grev. (= Odontella sinensis (Grev.)

Grun.)18,P1.25,Fig. 3
corpulenta Grev. 19, PI. 22, Fig. 6
?decorataGrev. 19, PI. 21, Fig. 8
elegantula Grev. 19, PI. 22, Figs 4, 5
fimbriata Grev. 1865 (=Odontella fimbriata (Grev.)

Schrader) 19, PI. 22, Figs 1,2

gigantea Grev. ( = Kittonia gigantea (Grev.) De

Toni) 19, PI. 23, Figs 1,2
inflata Grev. 19, PI. 23, Figs 3, 4
johnsoniana Grev. (=Cerataulus johnsonianus

(Grev.) Cleve) 19, PI. 22, Fig. 3; PI. 23, Figs 5, 6
longicruris Grev. 19, PI. 23, Figs 7-9
?mammosa Grev. 20, PI. 24, Figs 1, 2
minutissima Grev. 20, PI. 24, Fig. 3
nitida Grev. 20, PI. 24, Figs 4, 6
Ipodagrosa Grev. (=Hemiaulus podagrosa (Grev.)

De Toni) 20, PI. 24, Figs 5, 8
punctata Grev. 20, PI. 24, Fig. 7
roperiana Grev. (=Biddulphia aurita var. obesa

(Kiitz.) Must.) 20, PI. 24, Figs 9, 10; PI. 25, Figs 1,

2

sinuata Grev. 20
spinosa Grev. 20, PI. 25, Fig. 4
tenuicornis Grev. ( = Odontropis tenuicornis (Grev.)

Grun.) 21, PI. 25, Figs 5, 6
Brightwellia elaborata Grev. 21, PI. 25, Figs 7, 10

johnsonii Ralfs ex Grev. 21, PI. 25, Figs 8, 9
Campylodiscus ambiguus Grev. (=C. latus Shadbolt)

21, PL 26, Figs 1,2
biangulatus Grev. 21, PI. 26, Figs 3, 4
browneanus Grev. 21
crebrecostatus Grev. 21

diplostictus G. Norman ex Grev. 21, PI. 26, Figs 5, 6
ecclesianus Grev. 21, PI. 26, Figs 7, 8
fenestratus Grev. 21, PI. 26, Fig. 9
humilis Grev. 22, PI. 26, Fig. 10
imperialis Grev. 22, PI. 26, Fig. 11
kittonianus Grev. 22, PI. 27, Figs 1,2
normanianus Grev. (=C. wallichianus var.

normanianus (Grev.) De Toni) 22
notatus Grev. 22
ornatus Grev. 22

robertsianus Grev. 22, PI. 27, Figs 3, 4
stellatus Grev. 22, PI. 27, Figs 5, 6
undulatus Grev. ( C. brightwellii Grun.) 22, PI. 26,

Fig. 12

wallichianus Grev. 22, PI. 27, Figs 7, 8
Cestodiscus johnsonianus Grev. (= Coscinodiscus

johnsonianus (Grev.) Rattray) 23, PI. 27, Figs 10,

11
(?) ovalis Grev. (=Coscinodiscus biconfusus Van

Land.) 23, PI. 27, Fig. 9
pulchellus Grev. 1866 non Coscinodiscus pulchellus

Grev. 1866 (=C. superba Hardman ex Rattray) 23
stokesianus Grev. (= Coscinodiscus stokesianus

(Grev.) Grun.) 23
Cladogramma conicum Grev. 23
Clavularia barbadensis Grev. 23, PI. 28, Figs 7-10
Cocconeis armata Grev. (= Mastogloia oamaruensis

Cleve) 23, PI. 28, Fig. 6
arraniensis Grev. 24, PI. 29, Figs 1-3
barbadensis Grev. 24, PI. 29, Figs 5, 6
coelata Arn. ex Grev. (=Diploneis campy lodiscus

(Grun.) Cleve) 24, PI. 29, Fig. 4
crebrestriata Grev. (=C. scutellum Ehrenb.) 24, PI.

29, Figs 7, 8

grantiana Grev. 24, PI. 29, Fig. 9
granulifera Grev. (=C. distans Greg.) 24, PI. 29, Fig.

10

inconspicua Grev. 24
naviculoides Grev. (= Mastogloia barbadensis

(Grev.) Cleve) 24, PI. 29, Figs 11, 12

68 D. M. WILLIAMS

punctatissima Grev. (= Mastogloia punctatissima

(Grev.) Richard) 24, PI. 29, Figs 13, 14
regalis Grev. (=Campyloneis grevillei var. regalis

(Grev.) Cleve) 25, PI. 30, Figs 6, 7
Coscinodiscus angulatus Grev. 25, PI. 30, Fig. 3
armatus Grev. 25, PI. 30, Figs 4, 5
barbadensis Grev. 25
biradiatus Grev. (=Azpeitia biradiata (Grev.) P. A.

Sims) 25, PI. 31, Fig. 1
elegans Grev. 25, PI. 31, Figs 2, 4
elegantulus Grev. (=Azpeitia elegantula (Grev.) P.

A. Sims) 25, PI. 31, Fig. 3
excavatus Grev. ex Ralfs 26, PI. 32, Figs 3, 6, 7
griseusGrev. 26, PI. 31, Fig. 6
lewisianus Grev. 26, PI. 31, Figs 7, 9
macraeanus Grev. 26, PI. 31, Figs 5, 8
moronensis Grev. ex Rattray 26, PI. 30, Figs 1, 2
mossianus Grev. 26, PI. 30, Figs 1, 2
obliquus Grev. ex Rattray 26, PI. 32, Figs 2, 4, 5
oblongus Grev. (= Craspedodiscus ellipticus (Grev.)

Gombos) 26, PI. 28, Figs 1-4
patellaeformis Grev. 27, PI. 31, Fig. 10
pulchellus Grev. 27, PI. 33, Fig. 1
robustus Grev. ( = Endictya robusta (Grev.) Hanna

& Grant) 27, PI. 28, Fig. 5
scintillans Grev. ( = Actinocyclus actinochilus

(Ehrenb.) Simonsen) 27
splendidus Grev. 27, PI. 33, Figs 2, 5, 6
symmetricus Grev. 27, PI. 34, Figs 6-8
tuberculatus Grev. ( = Azpeitia tuberculata (Grev.)

P. A. Sims) 27, PI. 34, Figs 1,2
Cosmiodiscus barbadensis Grev. 27, PI. 34, Figs 3-5
elegans Grev. (= Coscinodiscus perikompsos

Rattray) 25, 27, PI. 34, Figs 3-4

normanianus Grev. (= Coscinodiscus apages Rattray) 27
Craspedodiscus umbonatus Grev. 28, PI. 33, Fig. 7
Craspedoporus johnsonianus Grev. 28

ralfsianus Grev. 28, PI. 34, Fig. 9
Creswellia annulata Grev. ( = Lauderia annulata

(Grev.) Cleve) 28, PI. 34, Figs 10, 1 1
barbadensis Grev. ( = Pyxidicula barbadensis (Grev.)

Strelnikova & Nikolaev) 28, PI. 35, Figs 1, 2
cylindracea Grev. 28
iferox Grev. (=P. ferox (Grev.) Strelnikova &

Nikolaev) 29, PI. 35, Figs 3, 4
minuta Grev. (=Stephanopyxis turris var. minuta

(Grev.) M. Perag.) 29, PI. 35, Fig. 5
palmeriana Grev. (= P. palmeriana (Grev.)

Strelnikova & Nikolaev) 29, PI. 35, Figs 7, 8
rudisGrev. 29, PI. 35, Fig. 6
sphaerica Grev. 29
superba Grev. 29, PI. 36, Figs 1-3
turgida Grev. 29, PI. 36, Figs 4-6
turris Grev. & Arn. (= P. turris (Grev.) Strelnikova

& Nikolaev) 30, PI. 36, Fig. 9
Cymatopleura angulata Grev. 30, PI. 37, Fig. 9
Cymbella lindsayana Grev. 30, PI. 37, Figs 7, 8
Diatoma brachygonium Grev. in Hook. 30
Diatomella balfouriana Grev. (-Grammatophora

balfouriana (Grev.) W. Sm.) 30, PI. 37, Figs 1,2
Dicladia? barbadensis Grev. ( = Dicladiopsis

barbadensis (Grev.) De Toni) 30, PI. 37, Fig. 3
Irobusta Grev. ( = Dicladiopsis barbadensis (Grev.)

De Toni) 30, 31, PI. 37, Fig. 6
Dictyopyxis brevis Grev. ( = Endictya oceanica

Ehrenb.) 31, PI. 37, Figs 4, 5

Echinella drcularis Grev. ( = Meridion circulare

(Grev.)Agardh)31
fasciculata [ft] truncata Grev. (= Synedra fasciculata

var. truncata (Grev.) Patr.) 31
Entogonia amabilis Grev. 31, PI. 37, Fig. 7
conspicua Grev. ( = Entogonia davyana var.

conspicua (Grev.) Bergon) 31
elegans Grev. 31
punctulata Grev. ( = E. pulcherrima var. punctulata

(Grev.) Bergon) 32
reticulata Grev. 32, PI. 37, Fig. 8
venulosa Grev. 32

Euodia barbadensis Grev. 32, PI. 37, Fig. 10
Eupodiscus barbadensis Grev. 32, PI. 38, Fig. 1
hardmanianus Grev. 32, Pi. 38, Figs 2, 3
jonesianusGrev. (= Coscinodiscus jonesianus

(Grev.) Ostenf.) 32, PI. 38, Fig 4-6
minutus Grev. 32
obscurus Grev. 32
oculatus Gfev. 33

parvulus Grev. ex Rattray 33, PI. 38, Figs 7, 8
punctulatus Grev. 33, PI. 38, Fig. 10
scaberGre\. (= Coscinodiscus scaber (Grev.)

Rattray) 33, PI. 38, Fig. 11
simplex Grev. 33, PI. 38, Fig. 9
trioculatus Grev. 33, PI. 39, Fig. 1
Exilaria fulgens Grev. (=Synedra fulgens (Grev.) W.

Sm.) 33

Fenestrella barbadensis Grev. 33, PI. 39, Fig. 2
Gephyria constricta Grev. ( = Entopyla australis

(Ehrenb.) Ehrenb.) 34, PI. 39, Fig. 3
gigantea Grev. (=Entopyla australis var. gigantea

(Grev.)Fricke)34

Glyphodesmis eximia Grev. 34, PI. 39, Figs 10, 11
Glyphodiscus stellatus Grev. 34, PI. 39, Figs 4, 5
Goniothecium prolongatum Grev. 34, PI. 39, Figs 6, 7
Grammatophora moronensis Grev. 34, PI. 39, Fig. 9
pusilla Grev. (=G. serpentina var. pusilla (Grev.)

VanHeurck)34,P1.39,Fig. 8
Heibergia barbadensis Grev. (= Entogonia barbadensis

(Grev.) Holmes & Brigger) 35, PI. 40, Figs 1-3
Heliopelta nitida Grev. (=Actinoptychus nitidus

(Grev.) Grundler) 35, PI. 40, Figs 4, 5
Hemiaulus alatus Grev. (=Riedelia alata (Grev.)

Schrader & Fenner) 35, PI. 40, Fig. 6
angustus Grev. 35, PI. 40, Figs 7-9
llcapitatus Grev. ( = Briggera capitata (Grev.) R.

Ross & P. A. Sims) 35, PI. 40, Figs 11, 12;
chinensis Grev. 35, PI. 41, Fig. 10
crenatus Grev. 35
exiguus Grev. 36, PI. 41 , Figs 1 , 2
hastatus Grev. 1, 35, 36, 61, PI. 41, Fig. 3
iobatus Grev. 36
longicornis Grev. (= Riedelia longicornis (Grev.)

Schrader & Fenner) 35, 36, PI. 40, Fig. 1.0
lyriformis Grev. (= Riedelia lyriformis (Grev.)

Schrader & Fenner) 35, 36, PI. 41, Fig. 4
minutus Grev. 36

mucronatus Grev. 36, PI. 41 , Figs 5-7
ornithocephalus Grev. ( = Briggera ornithocephala
(Grev.) R. Ross & P. A. Sims) 35, 36, PI. 41, Figs
8,9

pulvinatus Grev. 36, PI. 42, Figs 1, 4
punctatus Grev. 36
reticulatus Grev. 37
??robustus Grev. 37, PI. 42, Fig. 2

GREVILLE DIATOM TYPES

69

Hemiaulus - cont.

symmetricus Grev. 37, PI. 42, Figs 6, 7
tenuicornis Grev. (= Riedelia tenuicornis (Grev.)

Schrader & Fenner) 35, 37, PI. 43, Figs 1 , 2
Heterodictyon rylandsianum Grev. ( = Asterolampra
marginata (Brightw.) Grev.) 37, PI. 42, Figs 3, 5
splendidum Grev. (= Brightwellia splendida (Grev.)

Rattray)37, PI. 42, Fig. 8
Liradiscus barbadensis Grev. 37, PI. 42, Fig. 9
ellipticus Grev. 37, PI. 42, Fig. 10
minutus Grev. 37, PI. 43, Fig. 3
ovalis Grev. 37, PI. 43, Fig. 4
Mastogloia capitata Grev. 38, PI. 43, Figs 5, 6, 9
macdonaldii Grev. 38
minuta Grev. 38, PI. 43, Figs 7, 8
MelosiracostataGrev. (= Skeletonema costatum

(Grev.) Cleve) 38, PI. 44, Figs 1,2
nobilis Grev. 38, PI. 43, Figs 10, 1 1
setosa Grev. 38, PI. 44, Figs 3, 4
Monogramma smithiana Grev. ( = Achnanthes inflata

var. smithiana (Grev.) Greg.) 38
Navicula approximata Grev. 38, PI. 44, Fig. 15
californica Grev. (=N. hennedeyi var. californica

(Grev.) Cleve) 38, PI. 44, Figs 13, 14
Icistella Grev. (=Cistula lorenziana (Grun.) Cleve)

39, PI. 44, Fig. 5

compacta Grev. 39, PI. 44, Fig. 10

diversa Grev. ( = Diploneis? weisflogii (A. Schmidt)

Cleve) 39, PI. 44, Figs 8, 9
egyptiaca Grev. (=Oestrupia powelli var. egyptiaca

(Grev.)Heiden)39, PI. 44, Figs 7, 11,12, 16
excavata Grev. ( = N. spectabilis var. excavata

(Grev.) Cleve) 39
forcipata Grev. 39, PI. 45, Fig. 1
gemmata Grev. ( = Diploneis gemmata (Grev.)

Cleve) 39, PI. 45, Figs 2, 3
gregoriana Grev. ( = Lyrella lyra (Ehrenb.)

Karajeva) 40
indica Grev. ( = N. clavata var. indica (Grev.) Cleve)

40, PI. 45, Figs 4-6

irrorata Grev. 40, PI. 45, Figs 7, 8

jamaicensis Grev. (=Dictyoneis jamaicensis (Grev.)

Cleve) 40, PI. 45, Figs 9, 10
johnsoniana Grev. (-Trachyneis johnsoniana

(Grev.) Cleve) 40, PI. 46, Figs 1,2
lewisiana Grev. (=Frickia lewisiana (Grev.) Heiden)

40, PI. 46, Figs 3, 5, 6
luxuriosa Grev. 40

lyra var. recta Grev. 40, PI. 46, Figs 11, 12

notabills Grev. (=Diploneis notabilis (Grev.) Cleve)

41, PI. 46, Fig. 4
nummularia Grev. 41, PI. 46, Fig. 7

polysticta Grev. (=N. baileyana Grun.) 41, PI. 46,

Figs 8, 9
rimosa Grev. (=N. sandriana Grun.) 41, PI. 47, Figs

1-3

robertsiana Grev. 41, PI. 46, Fig. 10
spectalissima Grev. (=Dictyoneis marginata var.

spectalissima (Grev.) Cleve) 41, PI. 47, Figs 4, 5
strangulata Grev. (=Dictyoneis marginata (Lewis)

Cleve) 41
sulcata Grev. (=Cymatoneis sulcata (Grev.) Cleve)

41, PI. 47, Figs 6, 7
zanzibarica Grev. 41, PI. 48, Fig. 1
Odontella hastata (Grev.) J. Fenner ex D. M. Williams

1,61

Odontodiscus barbadensis Grev. 41, PI. 47, Fig. 8
Omphalopelta moronensis Grev. ( = Actinoptychus
moronensis (Grev.) Cleve) 41, PI. 48, Figs 8, 9
Omphalopsis australis Grev. 42, PI. 48, Fig. 2
Palmeria hardmaniana Grev. (=Hemidiscus
hardmanianus (Grev.) Kuntze) 42, PI. 48,
Figs 4-6

Peponia barbadensis Grev. 42, PI. 48, Fig. 7
Perissonoe cruciata (Janisch & Rabenh.) G. W.
Andrews & Stoelzel 61

parvula (Grev.) D. M. Williams 1 , 7, 61
Pinnularia hartleyana Grev. 42, PI. 48, Fig. 3
Plagiogramma? angulatum Grev. 42, PI. 49, Figs 1-3

atomus Grev. 42, PI. 49, Figs 4, 5

barbadense Grev. 42

californicum Grev. 42, PI. 49, Figs 9, 10

constrictum Grev. 42, PI. 49, Figs 6, 7

costatum Grev. 43, PI. 49, Fig. 8

decussatum Grev. 43

elongatum Grev. 43, PI. 49, Fig. 1 1

inaequale Grev. 43, PI. 50, Figs 1-3

jamaicense Grev. 43, PI. 49, Figs 12, 13

lyratum Grev. 43, PI. 50, Figs 4, 5

obesum Grev. 43, PI. 50, Fig. 10

orientale Grev. 43, PI. 50, Figs 8,11

ornatum Grev. 43, PI. 50, Fig. 9

pulchellum Grev. 44, PI. 50, Fig. 6

pygmaeum Grev. ( = P. pulchellum var. pygmaeum
(Grev.) H. & M. Perag.) 44, PI. 51, Figs 1,6

robertsianum Grev. 44

spectabile Grev. 44, PI. 51, Fig. 2

?tessellatum Grev. 44, PI. 50, Fig. 7

validum Grev. 44, PI. 51, Figs 3, 4

wallichianum Grev. 44, PI. 51, Figs 5, 7
Pleurosigma compactum Grev. ( = Gyrosigma

compactum (Grev.) Cleve) 44, PI. 51, Figs 8-10
Porodiscus concinus Grev. 44, PI. 52, Figs 1-3

elegans Grev. 45, PI. 52, Fig. 4

major Grev. 45, PI. 52, Figs 5, 6

nitidus Grev. 45, PI. 52, Figs 7, 8

oblongus Grev. (= Coscinodiscus oblongus (Grev.)
A. Schmidt) 45, PI. 53, Figs 2, 3, 5

splendidus Grev. (= Coscinodiscus splendidus

(Grev.) Gombos) 45, PI. 52, Fig. 9
Porpeia ornata Grev. ( = P. quadriceps var. ornata
(Grev.) F. W. Mills) 45, PI. 51 , Figs 11,12

quadrata Grev. 45, PL 53, Fig. 1
Pseudauliscus grevillei R. Ross 1, 62

nebulosus (Grev.) Rattray 62

nebulosus Leuduger-Fortmorel 62
Pyxilla barbadensis Grev. 45, PI. 53, Fig. 7

johnsoniana Grev. 46, PI. 53, Fig. 6
Rhaphoneis ampiceros var. cruciata (Janisch &
Rabenh.) Mereschk. 61

amphiceros var. tetragona Grun. 61
Rhizosolenia striata Grev. 46
Rutilaria elliptica Grev. 46, PI. 53, Fig. 4

epsilon Grev. 46, PI. 53, Fig. 10

obesa Grev. 46, PI. 53, Figs 8, 9

superba Grev. 46, PI. 53, Fig. 11

ventricosa Grev. 46

Rylandsia biradiata Grev. 46, PI. 54, Fig. 1
Skeletonema barbadense Grev. 47, PI. 54, Figs 8, 9
Spatangidium ralfsianum G. Norman ex Grev.

(=Asteromphalus heptactis (Breb.) Ralfs) 47, PI.
55, Figs 1-3

70 D. M. WILLIAMS

Stauroneis apiculata Grev. ( = Cocconeis apiculata

(Grev.) A. Schmidt) 47, PI. 54, Figs 2, 3
australis Grev. (= Navicula biformis (Grun.)

A. Mann) 47, PI. 54, Fig. 7
decora Grev. 47, PI. 54, Fig. 4
(?) obesa Grev. 47, PI. 54, Figs 5, 6
rotundata Grev. 47, PL 55, Fig. 4
scaphulaeformis Grev. 47
Stictodesmis australis Grev. (=Climaconeis lorenziana

Grun.) 47, PI. 55, Figs 5-7
Stictodiscus buryanus Grev. 48, PI. 55, Fig. 9; PI. 56,

Fig. 1

californicus Grev. 48, PI. 55, Fig. 8; PI. 56, Fig. 2
hardmanianus Grev. 48, PI. 56, Fig. 4
insignis Grev. 48, PI. 55, Fig. 10
johnsonianus Grev. 48, PI. 56, Fig. 3
kittonianus Grev. 48

Strangulonema barbadense Grev. 48, PI. 56, Figs 5, 6
Surirella eximia Grev. 48, PI. 56, Fig. 8
macraeana Grev. 48, PI. 56, Fig. 7
palmeriana Grev. 48, PI. 56, Fig. 9
superba Grev. 49, PI. 57, Fig. 1
Syndetocystis grevilleanus Walker & Chase (=5.

barbadensis Van Heurck) 49
Synedra clavata Grev. (=Synedrosphenia clavata

(Grev.) H. & M. Perag.) 49, PI. 57, Fig. 2
normaniana Grev. (=Licmophora normanianus

(Grev.) Wahrer) 49, PI. 57, Figs 3-5
Syringidium daemon Grev. (=Cerataulina bicorne

(Ehrenb.)Hasle)49

Terebraria barbadensis Grev. 50, PI. 57, Figs 6, 7
Thaumatonema barbadense Grev. 50, PI. 57, Fig. 9

?costatum Grev. 50, PI. 57, Fig. 8
Triceratium abercrombieanum Grev. ( = Entogonia
abercrombieana (Grev.) Grev.) 50, PI. 58, Figs 1,
2

acceptum Grev. 50, PI. 58, Fig. 3
aculeatum Grev. 50, PI. 58, Fig. 4
acutangulum Grev. 51, PI. 58, Figs 5, 8
amoenum Grev. 51, PI. 58, Figs 9, 10
antiquum Pant. 61
approximatum Grev. ( = Entogonia decora (Bergon)

Holmes & Brigger) 51, PI. 58, Fig. 11
araneosum Grev. 51, PI. 59, Fig. 1
areolatum Grev. 51, PI. 59, Fig. 2
atomus Grev. 51, PI. 59, Fig. 3
attenuatum Grev. 51, PI. 58, Figs 6, 7
barbadense Grev. 51
blanditum Grev. 51, PI. 59, Fig. 4
brevinervum Grev. (=T. tabellarium Brightwell) 51,

PI. 59, Fig. 5
browneanum Grev. (=T. reticulum Ehrenb.) 51, PI.

59, Fig. 6

cancellatum Grev. 52, PI. 59, Fig. 7
capitatum Ralfs ex Grev. 52, PI. 59, Figs 8, 9
cellulosum Grev. 52

cinnamomeum Grev. 52, PI. 59, Figs 10, 11
concinnum Grev. 52, PI. 59, Fig. 12
constans Grev. 52, PI. 60, Fig. 3
convexum Grev. 52
cornutum Grev. 52, PI. 60, Fig. 4
davyanum Grev. (= Entogonia davyanum (Grev.)

Grev.) 52, PI. 60, Fig. 5
decorum Grev. 53, PI. 60, Figs 7, 8
definitum Grev. 53
delicatum Grev. 53, PI. 60, Figs 6, 9

denticulatum Grev. 53, PI. 60, Fig 10

disciforme Grev. 53, PI. 60, Fig. 1 1

dobreeanum G. Norman ex Grev. 53, PI. 61, Figs 1, 2

dulce Grev. 53, PI. 60, Fig. 12

exornatum Grev. 53, PI. 61, Figs 4-6

figuratum Grev. 53, PI. 61 , Fig. 7

firmum Grev. (=T. arcticum Brightwell) 53, PI. 61,

Fig. 8

flexuosum Grev. 53, PI. 61 , Fig. 3
foveatum Grev. 53, PI. 61, Fig. 9
giganteum Grev. 54, PI. 62, Fig. 1
gratiosum Grev. ( = Entogonia gratiosa (Grev.)

Grev.) 54, PI. 62, Figs 2, 6
hardmanianum Grev. ( T. radiatum Brightw.) 54,

PI. 62, Figs 3-5, 7
harrisonianum G. Norman & Grev. ( = Stictodiscus

harrisonianus (Norm. & Grev.) Castr.) 54
implicitum Grev. 54, PI. 62, Figs 8, 9
inaequale Grev. 54, PI. 63, Fig. 1
inconspicuum Grev. (=Lisitzinia inconspicua

(Grev.) Gleser) 54, PI. 63, Figs 2, 3
inelegans Grev. 54, PI. 63, Figs 4, 7
inflatum Grev. (= Stictodiscus inflatum (Grev.)

Castr.) 54, PI. 63, Fig. 5
inglorium Grev. 54
inopinatum Grev. (=Entogonia inopinatum (Grev.)

Grev.) 54, PI. 63, Fig. 6
inornatum Grev. (=Pseudotriceratium wallichii

(Ralfs) Simonsen) 54, PI. 63, Fig. 8
insignis Grev. 55, PI. 64, Fig. 1
irregulare Grev. 55
kittonianum Grev. (=T? solenoceros (Ehrenb.) Van

Land.) 55, PI. 67, Fig. 3
labrynthaeum Grev. 55
latum Grev. 55

lautum Grev. 55, PI. 64, Figs 2, 4-6
ligulatum Grev. 55, PI. 64, Fig. 3
lineatum Grev. 55, PI. 64, Fig. 7
lineolatum Grev. 55, PI. 64, Figs 8, 9
lobatum Grev. 55

mammosum Grev. 55, PI. 64, Fig. 10
microcephalum Grev. 56, PI. 64, Fig. 11
microstictum Grev. 56, PI. 65, Figs 1-3
modestum Grev. 56, PI. 67, Fig. 5
moronense Grev. 56, PI. 65, Figs 4, 5
nebulosum Grev. 56
neglectum Grev. 56, PI. 65, Figs 7, 8
nitescens Grev. 56, PI. 65, Figs 10, 11
nitidum Grev. 56, PI. 66, Figs 1,2
normanianum Grev. 56
notabilis Grev. 56, PI. 67, Figs 1, 2, 4
obesum Grev. 56, PI. 65, Figs 6, 9
obscurum Grev. 57, 61, PI. 66, Fig. 3
oculatum Grev. 57, PI. 66, Figs 4, 5
ornatum Grev. 57, PI. 67, Fig. 6
pallidum Grev. 57, PI. 66, Figs 6, 7
partitum Grev. 57, PI. 66, Fig. 8
pauperculum Grev. 57, PI. 66, Fig. 9
pectinatum Grev. 57
perminutum Grev. (=Trinacria praetenue var.

perminuta (Grev.) Grun.) 57, PI. 68, Figs 1, 2
perpusillum Grev. 57, PI. 68, Figs 3, 6
picturatum Grev. 57, PI. 68, Figs 7, 8
plumosum Grev. 57, PI. 69, Figs 1, 4
polygonium Grev. (= Stictodiscus polygonius

(Grev.) Castr.) 58, PI. 69, Figs 2, 5

GREVILLE DIATOM TYPES

71

Triceratium - cont.
praetenue Grev. 58, PI. 68, Figs 4, 5
productum Grev. 58, PI. 69, Fig. 3
prominens Grev. 58
pulcherrimum Grev. ( = Entogonia pulcherrima

(Grev.) Grev.) 58, PI. 69, Figs 6, 7
quadrangulare Grev. (=T. formosum f.

quadrangulare (Grev.) Hust.) 58, PI. 70, Figs 7, 8
quadratum Grev. 58, PI. 70, Figs 2, 3
quadricorne Grev. (= Stictodiscus quadricornis

(Grev.) Castr.) 58, PI. 70, Figs 4, 5
quinquelobatum Grev. (=T. formosum f.

quinquelobatum (Grev.) Hust.) 58
repletum Grev. 58, PI. 71, Figs 1,2
reticulatum Grev. (= Stictodiscus? reticulcellulosus

Bergon & Tempere) 59, PI. 71, Figs 3, 4
robertsianum Grev. 59, PI. 71, Figs 5, 6
robertsianum Grev. 1866 (non T. robertsianum

Grev. 1863) 59, PI. 72, Figs 1,2
robustum Grev. 59, PI. 72, Figs 5, 6

roperianum Grev. 59, PI. 72, Figs 7, 8
rotundatum Grev. 59, PI. 71, Fig. 7
rylandsianum Grev. 59, PI. 72, Figs 3, 4
sexangulatum Grev. 59, PI. 73, Figs 1 , 2
smithianum Grev. 59, PI. 70, Fig. 6
stokesianum Grev. 59
subcapitatum Grev. 59, PI. 73, Figs 3, 4
tesselatum Grev. 59, PI. 73, Figs 5, 6
thwaitesianum Grev. 60, PI. 73, Figs 7, 8
trilineatum Grev. 60, PI. 73, Figs 9, 10
tumidum Grev. 60, PI. 73, Figs 11, 12
unguiculatum Grev. 60, PI. 74, Figs 1-3
variegatum Grev. (=Entogonia variegata (Grev.)

Grev.) 60

venulosum Grev. (=T. tabellarium Brightw.) 60
westianum Grev. 60, PI. 74, Fig. 5
zonatulatum Grev. 60, PI. 74, Figs 7, 8
zonatum Grev. 60, PI. 74, Fig. 4
Xanthiopyxis? umbonatus Grev. 61, PI. 74,

Figs 9-11

72 D. M. WILLIAMS

List of plates

Plate 1. Fig. 1 : Achnanthes angustata. Figs 2-4: A. carmichaelii. Figs 5-7: A. pennaeformis. Figs 8-10: A.

costata. Figs 11-12: A. gregoriana.
Plate 2. Fig. \\Actinodiscus barbadensis. Figs 2, 4, 5: Amphiprora brebissoniana. Fig. 3: A. hyalina. Fig.

6: A chinensis. Fig. 7: A jolisiana. Fig. 8: A clepsydra.
Plate 3. Figs 1-2: Amphiprora conspicua. Figs 3-5: A delicatula. Fig. 6: A oblonga. Fig. 7: A eximia.

Figs 8-9: A /epufa.
Plate 4. Figs 1-2: Amphiprora kuetzingiana. Figs 3-4: A /ata. Figs 5-6: A lineata. Figs 7-8: /I.

meneg/z/m'ana.
Plate 5. Figs 1-2: Amphiprora nitida. Figs 3-5: A oblonga. Fig. 6: A rectangularis. Fig. 7: A paradoxa.

Fig. 8: A rabenhorstiana. Figs 9-10: A thwaitesiana.
Plate 6. Figs 1-3: Amphiprora superba. Fig. 4: Amphitetras elegans. Fig. 5: A minuta. Figs 6-7: A

parvula. Fig. 8: A elegans. Fig. 9: A producta. Fig. 10: A punctata.
Plate 7. Figs 1-2: Amphitetras radiata. Fig. 3: Amphora undulata. Fig. 4: A sinuata. Fig. 5: A.flexuosa.

Figs 6-7: A pulchra. Figs 8-9: A magnifica.
Plate 8. Figs 1-2: Asterolampra aemulans. Figs 3-6: A 'affinis.
Plate 9. Fig. 1: Asterolampra aliena. Fig 2: A ambigua var. ft. Figs 3-4: A vanabilis. Fig. 5: A concinna.

Fig. 6: A crenata. Fig. 7: A concinna.

Plate 10. Figs 1-4: Asterolampra decora. Figs 5-8: A decorata. Figs 9-10: A eximia.
Plate 11. Figs 1-2: Asterolampra hiltoniana. Fig. 3: A kittoniana. Figs 4-5: A /aevw. Figs 6-7: A

pulchra. Figs 8-9: A punctata.
Plate 12. Fig. 1: Asterolampra ralfsiana. Fig. 2: A roperiana. Fig. 3: A rylandsiana. Fig. 4: A scutula.

Fig. 5: A shadboltiana. Fig. 6: A simulans. Fig. 7: /I. stellulata. Figs 8-12: A vanabilis.
Plate 13. Figs 1-2: Asterolampra vanabilis. Figs 3-9: A vulgaris var. a. Fig. 10: A vulgaris var. ft. Figs

11-12: A vulgaris var. c.
Plate 14. Figs 1-2: Aulacodiscus angulatus. Fig. 3: A amoenus. Fig. 4: Asteromphalus elegans. Fig. 5:

Asterolampra wallichiana. Fig. 6: A vulgaris var. e.
Plate 15. Figs 1-2: Aulacodiscus extans. Fig. 3: A jonesianus. Fig. 4: A gigos. Fig. 5: A decorus. Fig. 6:

A cinctus. Figs 7-8: A decorus. Fig. 9: A cinctus. Figs 10-11: A attenuatus.
Plate 16. Figs 1-2: Aulacodiscus inflatus. Fig. 3: A macreanus. Fig. 4: A inflatus. Fig. 5: A kilkellyanus.

Fig. 6: A paradoxus. Figs 7-8: A mammosus.
Plate 17. Figs 1-3: Aulacodiscus orientalis. Figs 4-5: A pellucidus. Fig. 6: A sparsus. Fig. 7: A

Plate 18. Figs 1-2. Aulacodiscus spectabilis. Figs 3-4: Auliscus ambiguus. Fig. 5: A barbadensis. Fig. 6: A

elaboratus. Figs 7-8: A cellulatus.
Plate 19. Fig. 1: Auliscus elegans. Fig. 2: A hardmanianus. Fig. 3: A johnsonianus. Fig. 4: A elegans.

Fig. 5: A johnsonianus. Fig. 6: A moronensis. Fig. 7: A macreanus. Fig. 8: A nebulosus.
Plate 20. Figs 1-2: A macreanus. Fig. 3: A normanianus. Fig. 4: A notatus. Fig. 5: A parvulus. Figs

6-7: A ornatus.
Plate 21. Fig. 1: Aufocws racemosus. Figs 2-3: A reticulatus. Figs 4-5: A racemosus. Fig. 6: A

ralfsianus. Fig. 7: Berkeley a fragilis. Fig. 8: Biddulphia decorata.
Plate 22. Figs 1-2: Biddulphia fimbriata. Fig. 3: 5. johnsoniana. Figs 4-5: 5. elegantula. Fig. 6: fi.

corpulenta. Figs 7-9: 5. barbadensis.
Plate 23. Figs 1-2: Biddulphia gigantea. Figs 3-4: 5. w/7ata. Figs 5-6: . johnsoniana. Figs 7-9: fl.

/ong/crarw.
Plate 24. Figs 1-2: Biddulphia mammosa. Fig. 3: fl. minutissima. Fig. 4: fi. m'ftWa. Fig. 5: 5. podagrosa.

Fig. 6: . nitida. Fig. 7: B. punctata. Fig. 8: #. podagrosa.
Plate 25. Figs 1-2: Biddulphia roperiana. Fig. 3: 5. chinensis. Fig. 4: 5. spinosa. Figs 5-6: 5. tenuicornis.

Fig. 7: Brightwellia elaborata. Figs 8-9: 5. johnsoni. Fig. 10: . elaborata.
Plate 26. Figs 1-2: Campylodiscus ambiguus. Figs 3-4: C. biangulatus. Figs 5-6: C. diplostictus. Figs

7-8: C. ecclesianus. Fig. 9: C. fenestratus. Fig. 10: C. humilis. Fig. 11: C. imperialis. Fig. 12: C.

Plate 27. Figs 1-2: Campylodiscus kittonianus. Figs 3-4: C. robertsianus. Figs 5-6: C. stellatus. Figs 7-8:

C. wallichianus. Fig. 9: Cestodiscus ovalis. Figs 10-11: C. johnsonianus.
Plate 28. Figs 1-4. Coscinodiscus oblongus. Fig. 5: C. robustus. Fig. 6: Cocconeis armata. Figs 7-10.

Clavularia barbadensis.
Plate 29. Figs 1-3: Cocconeis arraniensis. Fig. 4: C. coelata. Figs 5-6: C. barbadensis. Figs 7-8: C.

GREVILLE DIATOM TYPES 73

crebrestriata. Fig. 9: C. grantiana. Fig. 10: C. granulifera. Figs 11-12: C. naviculoides. Figs 13-14: C.

Plate 30. Figs 1-2: Coscinodiscus moronensis. Fig. 3: C. angulatus. Figs 4-5: C. armatus. Figs 6-7:

Cocconeis regalis.
Plate 31. Fig. 1 : Coscinodiscus biradiatus. Fig. 2: C. elegans. Fig. 3: C. elegantulus. Fig. 4: C. elegans. Fig.

5: C. macreanus. Fig. 6: C. griseus. Fig. 7: C. lewisianus. Fig. 8: C. macreanus. Fig. 9: C. lewisianus. Fig.

10: C. patellaeformis.
Plate 32. Fig. 1 : Coscinodiscus mossianus. Fig. 2: C. obliguus. Fig. 3: C. excovatas. Figs 4-5: C. obliquus.

Figs 6-7: C. excavatas.
Plate 33. Fig. 1: Coscinodiscus pulchellus. Fig. 2: C. splendidus. Figs 3-4: C. elegans. Figs 5-6: C.

splendidus. Fig. 7: Craspedodiscus umbonatus.
Plate 34. Figs 1-2: Coscinodiscus tuberculatus. Figs 3-5: Cosmiodiscus barbadensis. Figs 6-8: Coscino-

discus symmetricus. Fig. 9: Craspedoporus ralfsianus. Figs 10-11: Creswellia annulata.
Plate 35. Figs 1-2: Creswellia barbadensis. Figs 3-4: C. /era*. Fig. 5: C. minuta. Fig. 6: C. radzs. Figs 7-8:

C. palmeriana.
Plate 36. Figs 1-3: Creswellia superba. Figs 4-6: C. turgida. Figs 7-8: Cymbella lindsayana. Fig. 9:

Creswellia turns.
Plate 37. Figs 1-2: Diatomella balfouriana. Fig. 3: Dicladia barbadensis. Figs 4-5: Dictyopyxis brevis.

Fig. 6: Dicladia robusta. Fig. 7: Entogonia amabilis. Fig. 8: E. reticulata. Fig. 9: Cymatopleura angulata.

Fig. 10: Euodia barbadensis.
Plate 38. Fig. 1: Eupodiscus barbadensis. Figs 2-3: E. hardmanianus. Figs 4-6: E. jonesianus. Figs 7-8:

E. parvulus. Fig. 9: . simplex. Fig. 10: E. punctulatus. Fig. 11: E. scaber.
Plate 39. Fig. 1: Eupodiscus trioculatus. Fig. 2: Fenestrella barbadensis. Fig. 3: Gephyria constricta. Figs

4-5: Glyphodiscus stellulatus. Figs 6-7: Goniothecium prolongatum. Fig. 8: Grammatophora pusilla.

Fig. 9: G. moronensis. Figs 10-11: Glyphodesmis eximia.
Plate 40. Figs 1-3: Heibergia barbadensis. Figs 4-5: Heliopelta nitida. Fig. 6: Hemiaulus alatus. Figs 7-9:

H. angustus. Fig. 10: //. longicornis. Figs 11-12: //. capitatus.
Plate 41. Figs 1-2: Hemiaulus exiguus. Fig. 3: //. hastatus. Fig. 4: //. lyriformis. Figs 5-7: //. mucronatus.

Figs 8-9: //. ornithocephalus. Fig. 10: //. chinensis.
Plate 42. Fig. 1: Hemiaulus pulvinatus. Fig. 2: //. robustus. Fig. 3: Heterodictyon rylandsianum. Fig. 4: H.

pulvinatus. Fig. 5: Heterodictyon rylandsianum. Figs 6-7: Hemiaulus symmetricus. Fig. 8: Heterodictyon

splendidum. Fig. 9: Liradiscus barbadensis. Fig. 10: L. ellipticus.
Plate 43. Figs 1-2: Hemiaulus tenuicornis. Fig. 3: Liradiscus minutus. Fig. 4: L. ovalis. Figs 5-6:

Mastogloia capitata. Figs 7-8: Af. minuta. Fig. 9: M. capitata. Figs 10-11: Melosira nobilis.
Plate 44. Figs 1-2: Melosira costata. Figs 3-4: M. setosa. Figs 5-6: Navicula cistella. Fig. 7. W. egyptiaca.

Figs 8-9: M diversa. Fig. 10: M compacta. Figs 11-12: JV. egyptiaca. Figs 13-14: M californica. Fig. 15:

M approximata. Fig. 16: TV. egyptiaca.
Plate 45. Fig. 1: Navicula forcipata. Figs 2-3: M gemmata. Figs 4-6: M indica. Figs 7-8: AT. irrorata. Figs

9-10: W. jamaicensis.
Plate 46. Figs 1-2: Navicula johnsoniana. Fig. 3: N. lewisiana. Fig. 4: M notabilis. Figs 5-6: M lewisiana.

Fig. 7: M nummularia. Figs 8-9: N. polysticta. Fig. 10: M robertsiana. Figs 11-12: N. /yra var. recto.
Plate 47. Figs 1-3: Navicula rimosa. Figs 4-5: M spectalissima. Figs 6-7: M sulcata. Fig. 8: Odontodiscus

barbadensis.
Plate 48. Fig. 1: Navicula zanzibarica. Fig. 2: Omphalopsis australis. Fig. 3: Pinnularia hartley ana. Figs

4-6: Palmeria hardmaniana. Fig. 7: Peponia barbadensis. Figs 8-9: Omphalopelta moronensis.
Plate 49. Figs 1-3: Plagiogramma angulatum. Figs 4-5: P. atomus. Figs 6-7: P. constrictum. Fig. 8: P.

costatum. Figs 9-10: P. californicum. Fig. 11: P. elongatum. Figs 12-13: P. jamaicense.
Plate 50. Figs 1-3: Plagiogramma inaequale. Figs 4-5: P. lyratum. Fig. 6: P. pulchellum. Fig. 7: P.

tessellatum. Fig. 8: P. orientale. Fig. 9: P. ornatum. Fig. 10: P. obesum. Fig. 11: P. orientale.
Plate 51. Fig. 1: Plagiogramma pygmaeum. Fig. 2: P. spectabile. Figs 3-4: P. validum. Fig. 5: P.

wallichianum. Fig. 6: P. pygmaeum. Fig. 7: P. wallichianum. Figs 8-10. Pleurosigma compactum. Figs

11-12: Porpeia ornata.
Plate 52. Figs 1-3: Porodiscus concinus. Fig. 4: P. elegans. Figs 5-6: P. mayor. Figs 7-8: P. rutidus. Fig. 9:

P. splendidus.
Plate 53. Fig. 1: Porpeia quadrata. Figs 2-3: Porodiscus oblongus. Fig. 4: Rutilaria elliptica. Fig. 5:

Porodiscus oblongus. Fig. 6: Pyxilla johnsoniana. Fig. 7: P. barbadensis. Figs 8-9: Rutilaria obesa. Fig.

10: /?. epsilon. Fig. 11: fl. superba.
Plate 54. Fig. 1 : Rylandsia biradiata. Figs 2-3: Stauroneis apiculata. Fig. 4: S. decora. Figs 5-6: 5.

Fig. 7: S. australis. Figs 8-9: Skeletonema barbadense.

74 D. M. WILLIAMS

Plate 55. Figs 1-3: Spatangidium ralfsianum. Fig. 4: Stauroneis rotundata. Figs 5-7: Stictodesmis

australis. Fig. 8: Stictodiscus calif ornicus . Fig. 9: S. buryanus. Fig. 10: 5. insignis.
Plate 56. Fig. 1: Stictodiscus buryanus. Fig. 2: 5. californicus. Fig. 3: 5. johnsonianus. Fig. 4: S.

hardmanianus. Fig. 5-6: Strangulonema barbadense. Fig. 7: Surirella macreana. Fig. 8: 5. eximia. Fig. 9:

5. palmeriana.
Plate 57. Fig. 1 : Surirella superba. Fig. 2: Synedra clavata. Figs 3-5: 5. normaniana. Figs 6-7: Terebraria

barbadensis. Fig. 8: Thaumatonema costatum. Fig. 9: T. barbadense.
Plate 58. Figs 1-2: Triceratium abercrombeanum. Fig. 3: T acceptum. Fig. 4: 7 1 . aculeatum. Fig. 5: T.

acutangulum. Figs 6-7: 7. attenuatum. Fig. 8: T. acutangulum. Figs 9-10: T. amoenum. Fig. 11: T.

approximatum .
Plate 59. Fig. 1: Triceratium araneosum. Fig. 2: 7". areolatum. Fig. 3: T. atomus. Fig. 4: T. blanditum. Fig.

5: 7 1 . brevinervum. Fig. 6: T. broweanum. Fig. 7: T cancellatum. Fig. 8-9: T. capitatum. Figs 10-11: 7 1 .

cinnamomeum. Fig. 12: T. concinnum.
Plate 60. Figs 1-2: Triceratium cinnamomeum. Fig. 3: T. constans. Fig. 4: T. cornutum. Fig. 5: T.

davyanum. Fig. 6: T. delicatum. Figs 7-8: 7". decorum. Fig. 9: 7. delicatum. Fig. 10: 7. denticulatum. Fig.

11: 7. disciformis. Fig. 12: 7. du/ce.
Plate 61. Figs 1-2: Triceratium dobreeanum. Fig. 3: 7. flexuosum. Figs 4-6: 7. exornatum. Fig. 7: 7.

figuratum. Fig. 8: T.firmum. Fig. 9: T.foveatum.
Plate 62. Fig. 1. Triceratium giganteum. Fig. 2: 7. gratiosum. Figs 3-5: 7. hardmanianum. Fig. 6: 7.

gratiosum. Fig. 7: 7. hardmanianum. Figs 8-9: 7. implicitum.
Plate 63. Fig. 1: Triceratium inaequale. Figs 2-3: T. inconspicuum. Fig. 4: 7. inelegans. Fig. 5: 7.

inflatum. Fig. 6: T. inopinatum. Fig. 7: 7. inelegans. Fig. 8: T. inornatum.
Plate 64. Fig. 1: Triceratium insignis. Fig. 2: 7. lautum. Fig. 3: T ligulatum. Figs 4-6: 7. lautum. Fig. 7: 7.

lineatum. Figs 8-9: 7. lineolatum. Fig. 10: 7. mammosum. Figs 11-12: 7. microcephalum.
Plate 65. Figs 1-3: Triceratium microstictum. Figs 4-5: 7. moronense. Fig. 6: 7. obesum. Figs 7-8: 7.

neglectum. Fig. 9: 7. obesum. Figs 10-11: 7. nitescens.
Plate 66. Figs 1-2: Triceratium nitidum. Fig. 3: 7. obscurum. Figs 4-5: 7. oculatum. Figs 6-7: T.

pallidum. Fig. 8: 7. partitum. Figs 9-10: 7. pauperculum.
Plate 67. Figs 1-2: Triceratium notabilis. Fig. 3: 7. kittonianum. Fig. 4: 7. notabilis. Fig. 5: 7. modestum.

Fig. 6: r. ornatum.
Plate 68. Figs 1-2: Triceratium perminutum. Fig. 3: 7. perpusillum. Figs 4-5: T. praetenue. Fig. 6: T.

perpusillum. Figs 7-8: T. picturatum.
Plate 69. Fig. 1: Triceratium plumosum. Fig. 2: 7. polygonium. Fig. 3: T. productum. Fig. 4: T.

plumosum. Fig. 5: T. polygonium. Figs 6-7: T. pulcherimum.
Plate 70. Fig. 1: Triceratium pulcherimum. Figs 2-3: 7 1 . quadratum. Figs 4-5: T. quadricorne. Fig. 6: T.

smithianum. Figs 7-8: T quadr angular e.
Plate 71. Figs 1-2: Triceratium repletum. Figs 3-4: T. reticulatum. Figs 5-6: 7. robertsianum 1863. Fig. 7:

r. rotundatum.
Plate 72. Figs 1-2: Triceratium roberstianum 1866. Figs 3-4: T. rylandsianum. Figs 5-6: T. robustum.

Figs 7-8: T. roperianum.
Plate 73. Figs 1-2: Triceratium sexangulatum. Figs 3-4: T. subcapitatum. Figs 5-6: T. tessellatum. Figs

7-8: r. thwaitesianum. Figs 9-10: T. trilineatum. Figs 11-12: T. tumidum.
Plate 74. Figs 1-3: Triceratium unguiculdtum. Fig. 4: 7". zonatum. Fig. 5: T. westianum. Figs 6-8: T.

zonatulatum. Figs 9-11: Xanthiopyxis umbonatus.

PLATE 1

75

76

PLATE 2

PLATE 3

77

78

PLATE 4

PLATE 5

79

80

PLATE 6

PLATE 7

81

82

PLATE 8

PLATE 9

83

84

PLATE 10

PLATE 1 1

85

86

PLATE 12

PLATE 13

87

PLATE 14

PLATE 1 5

89

90

PLATE 16

PLATE 17

91

92

PLATE 18

PLATE 19

93

r IP

J~-
s

94

PLATE 20

PLATE 21

95

PLATE 23

97

98

PLATE 24

100

PLATE 26

PLATE 27

101

102

PLATE 28

PLATE 29

103

104

PLATE 30

PLATE 3 1

105

106

PLATE 32

PLATE 33

107

108

PLATE 34

H

PLATE 35

109

110

PLATE 36

PLATE 37

111

112

PLATE 38

PLATE 39

113

PLATE 41

115

116

PLATE 42

PLATE 43

117

118

PLATE 44

PLATE 45

119

PLATE 47

121

122

PLATE 48

PLATE 49

123

PLATE 5 1

125

1

126

PLATE 52

128

PLATE 54

PLATE 55

129

130

PLATE 56

PLATE 57

131

132

PLATE 58

PLATE 59

133

134

PLATE 60

PLATE 61

135

136

PLATE 62

PLATE 63

137

138

PLATE 64

PLATE 65

139

140

PLATE 66

PLATE 67

141

142

PLATE 68

PLATE 69

143

144

PLATE 70

PLATE 71

145

146

PLATE 72

PLATE 73

147

148

PLATE 74

British Museum (Natural History)

MACROLICHENS OF EAST AFRICA

T. D. V. Swinscow & H. Krog

DrSwinscow was formerly Deputy Editor of the British Medical Journal.
Dr Krog is Professor of Taxonomic Botany at the University of Oslo.

This book is based mainly on collections made in the field by the authors. It covers
77 genera and 629 species. It is the first substantial study of a tropical lichen flora to
be undertaken by modern research methods. Thin-layer chromatography has been
used throughout, and the great majority of species have been studied by
microscopic examination of microtome sections. The nomenclature has been
thoroughly revised, and in all cases the basionym is given. The book will be
indispensable to students of the lichens of the African continent and valuable to
readers interested in lichens throughout the tropics.

Summer 1988, viii + 384pp, 185 figs., 16pp colour illustrations.
Hardback. 565 01039 5. 40.00

Photoset by Rowland Phototypesetting Ltd, Bury St Edmunds, Suffolk
Printed in Great Britain by Henry Ling Ltd, Dorchester